무)고현숙_국문안표지

제 권 호 게류와조에아유생 II 절지동물문 : 연갑강 : 십각목 : 단미하목 : 성산부채게상과, 애기털보부채게상과, 부채게상과 환경부국립생물자원관

제 권 호게류와조에아유생 II 절지동물문 : 연갑강 : 십각목 : 단미하목 : 성산부채게상과, 애기털보부채게상과, 부채게상과 집필자 : 고현숙, 이석현 신라대학교

,,,,. :

2010. 21.,... 4 4 7,709 65 216 World monograph 2, 1,313 25.,..,.

1 3 6 7 11 1. Eriphia smithii MacLeay 12 2. Hypothalassia armata (De Haan) 14 3. Sphaerozius nitidus Stimpson 15 4. Halimede fragifer (De Haan) 18 5. Parapanope euagora De Man 19 6. Echinoecus nipponicus Miyake 21 7. Harrovia elegans De Man 22 8. Harrovia japonica Balss 22 9. Actumnus elegans De Man 24 10. Actumnus marissinicus Takeda and Kim 24 11. Benthopanope indica (De Man) 25 12. Heteropilumnus ciliatus (Stimpson) 26 13. Neoactumnus convexus T. Sakai 27 14. Pilumnopeus granulatus Balss 28 15. Pilumnopeus makianus (Rathbun) 29 16. Pilumnus longicornis Hilgendorf 30 17. Pilumnus minutus De Haan 31 18. Pilumnus trispinosus (T. Sakai) 31 19. Zehntneriana amakusae (Takeda and Miyake) 33 20. Actaea polyacantha (Heller) 36 21. Actaea semblatae Guinot 37 22. Actaeodes hirsutissimus (Rüppell) 38 23. Forestia depressa (White) 39 24. Gaillardiellus orientalis (Odhner) 40 25. Novactaea pulchella (A. Milne-Edwards) 41 26. Pilodius miersi (Ward) 43 27. Pilodius nigrocrinitus Stimpson 43 28. Medaeops granulosus (Haswell) 45

2 II 29. Palapedia integra (De Haan) 46 30. Liomera margaritata (A. Milne-Edwards) 47 31. Neoliomera insularis (Adams and White) 48 32. Lybia caestifera (Alcock) 49 33. Cycloxanthops truncatus (De Haan) 50 34. Leptodius exaratus (H. Milne-Edwards) 51 35. Macromedaeus distinguendus (De Haan) 52 36. Microcassiope orientalis Takeda and Miyake 53 37. Nanocassiope granulipes (T. Sakai) 54 38. Neoxanthops lineatus (A. Milne-Edwards) 56 39. Banareia subglobosa (Stimpson) 56 40. Calvactaea tumida Ward 57 41. Atergatis floridus (Linnaeus) 59 42. Atergatis reticulatus (De Haan) 60 43. Atergatopsis germaini A. Milne-Edwards 61 44. Platypodia tomentosa (De Man) 62 63 69 93 95

3 Subphylum Crustacea Brünnich, 1772 Class Malacostraca Latreille, 1802 Subclass Eumalacostraca Grobben, 1892 Order Decapoda Latreille, 1802 Infraorder Brachyura Latreille, 1802 Superfamily Eriphioidea MacLeay, 1838 Family Eriphiidae MacLeay, 1838 Genus Eriphia Latreille, 1817 Eriphia smithii MacLeay, 1838 Family Hypothalassiidae Karasawa and Schweitzer, 2006 Genus Hypothalassia Gistel, 1848 Hypothalassia armata (De Haan, 1835) Family Menippidae Ortmann, 1893 Genus Sphaerozius Stimpson, 1858 Sphaerozius nitidus Stimpson, 1858 Superfamily Pilumnoidea Samouelle, 1819 Family Galenidae Alcock, 1898 Genus Halimede De Haan, 1833 Halimede fragifer (De Haan, 1835) Genus Parapanope De Man, 1895 Parapanope euagora De Man, 1895 Family Pilumnidae Samouelle, 1819 Genus Echinoecus Rathbun, 1894 Echinoecus nipponicus Miyake, 1939 Genus Harrovia Adams and White, 1849 Harrovia elegans De Man, 1887 Harrovia japonica Balss, 1921 Genus Actumnus Dana, 1851 Actumnus elegans De Man, 1887 Actumnus marissinicus Takeda and Kim, 1977 Genus Benthopanope Davie, 1989 Benthopanope indica (De Man, 1887)

4 II Genus Heteropilumnus De Man, 1895 Heteropilumnus ciliatus (Stimpson, 1858) Genus Neoactumnus T. Sakai, 1965 Neoactumnus convexus T. Sakai, 1965 Genus Pilumnopeus A. Milne-Edwards, 1867 Pilumnopeus granulatus Balss, 1933 Pilumnopeus makianus (Rathbun, 1931) Genus Pilumnus Leach, 1815 Pilumnus longicornis Hilgendorf, 1878 Pilumnus minutus De Haan, 1835 Pilumnus trispinosus (T. Sakai, 1965) Genus Zehntneriana Ng and Takeda, 2010 Zehntneriana amakusae (Takeda and Miyake, 1969) Superfamily Xanthoidea MacLeay, 1838 Family Xanthidae MacLeay, 1838 Genus Actaea De Haan, 1833 Actaea polyacantha (Heller, 1861) Actaea semblatae Guinot, 1976 Genus Actaeodes Dana, 1852 Actaeodes hirsutissimus (Rüppell, 1830) Genus Forestia Guinot, 1976 Forestia depressa (White, 1848) Genus Gaillardiellus Guinot, 1976 Gaillardiellus orientalis (Odhner, 1925) Genus Novactaea Guinot, 1976 Novactaea pulchella (A. Milne-Edwards, 1865) Genus Pilodius Dana, 1851 Pilodius miersi (Ward, 1936) Pilodius nigrocrinitus Stimpson, 1859 Genus Medaeops Guinot, 1967 Medaeops granulosus (Haswell, 1882) Genus Palapedia Ng, 1993 Palapedia integra (De Haan, 1835) Genus Liomera Dana, 1851 Liomera margaritata (A. Milne-Edwards, 1873) Genus Neoliomera Odhner, 1925 Neoliomera insularis (Adams and White, 1849)

5 Genus Lybia H. Milne Edwards, 1834 Lybia caestifera (Alcock, 1898) Genus Cycloxanthops Rathbun, 1897 Cycloxanthops truncatus (De Haan, 1837) Genus Leptodius A. Milne-Edwards, 1863 Leptodius exaratus (H. Milne-Edwards, 1834) Genus Macromedaeus Ward, 1942 Macromedaeus distinguendus (De Haan, 1835) Genus Microcassiope Guinot, 1967 Microcassiope orientalis Takeda and Miyake, 1969 Genus Nanocassiope Guinot, 1967 Nanocassiope granulipes (T. Sakai, 1939) Genus Neoxanthops Guinot, 1968 Neoxanthops lineatus (A. Milne-Edwards, 1867) Genus Banareia A. Milne-Edwards, 1869 Banareia subglobosa (Stimpson, 1858) Genus Calvactaea Ward, 1933 Calvactaea tumida Ward, 1933 Genus Atergatis De Haan, 1833 Atergatis floridus (Linnaeus, 1767) Atergatis reticulatus (De Haan, 1835) Genus Atergatopsis A. Milne-Edwards, 1862 Atergatopsis germaini A. Milne-Edwards, 1865 Genus Platypodia Bell, 1835 Platypodia tomentosa (De Man, 1902)

6,. 30 50 m,... 1., 7, 3 5. 1, 2, 1, 2 ( 2B, C). Kim (1973) (Eriphioidea) 2 ( Hypothalassia armata, Sphaerozius nitidus), (Pilumnoidea) 7 ( Harrovia elegans, Actumnus marissinicus, Benthopanope indica, Heteropilumnus ciliatus, Pilumnopeus makianus, Pilumnus longicornis, P. minutus), (Xanthoidea) 9 ( Actaea semblatae, Gaillardiellus orientalis, Medaeops granulosus, Neoliomera insularis, Leptodius exaratus, Macromedaeus distinguendus, Banareia subglobosa, Atergatis floridus, A. reticulatus), 18, 44., (zoea).,. 1, 2 ( 1 4 ) 2....,.,.,, 15... 1.

7. Lecia EZ40, Olympus E-30. (CL) (CW). Serène (1984). Leitz zoom Leitz Laborlux S. Leitz photoshop. 95%. (CL), (RDL)., (Clark et al., 1998). Ng (2008). (superfamily) (family). (NIBR). 1 2. 5 4 ( 1). ( 2A). 3 ( 1B, 2E). 2 ( ) ( 2B D), 4. 3 (,, )... 3, 1, 2, 1, 2 ( 3).

8 II 집게다리 걷는다리 A B 1. ( Leptodius exaratus). A. ; B..

9 앞옆구역 A 뒤옆구역 제 1 생식지 B 제 2 생식지 C 입안마루 제 1 생식지 제 2 생식지 D E 2.. A. ( Leptodius exaratus); B. ( Eriphia smithii); C. ( Pilumnus minutus); D. ( Leptodius exaratus); E. ( Pilumnus minutus).

10 II 촉모 갑각등가시 B 강모 갑각옆가시 C 외지 ( 바깥다리 ) 원지 갑각 복부 제 2 악지 D 제 1 촉각 제 1 악지 꼬리마디 내지 ( 안다리 ) 갑각이마가시 A 제 2 촉각 기절내돌기 저절내돌기 옆돌기 E F 기절 ( 밑마디 ) 외지 ( 바깥다리 ) 옆가시 내지 ( 안다리 ) 등가운데가시 I 포크 H G 3. ( Palapedia integra). A. ; B. 1 ; C. 2 ; D. ; E. 1 ; F. 2 ; G. 1 ; H. 2 ; I..

11 Subphylum Crustacea Brünnich, 1772 Class Malacostraca Latreille, 1802 Subclass Eumalacostraca Grobben, 1892 Order Decapoda Latreille, 1802 Infraorder Brachyura Latreille, 1802 Superfamily Eriphioidea MacLeay, 1838. 7. 1 2 1 ( 2B). 1. 2 Menippidae 2. Eriphiidae Hypothalassiidae

12 II 1.. 2 1/3 1 3. 1, 2, 1, 2+4 3+2+3. 2 5 Eriphia smithii. 2 4/5 1 1. 1, 2, 1, 4 3+3. 2, 3 Sphaerozius nitidus Family Eriphiidae MacLeay, 1838..... : 2 ( 1 ), 8 ( 1 ). :. Genus Eriphia Latreille, 1817.. 2.. : 7 ( 1 ). 1. Eriphia smithii MacLeay, 1838 ( 1A, B) Eriphia smithii MacLeay, 1838, p. 60; Takeda, 1982, p. 184, fig. 544. Eriphia sebana smithii: Stimpson, 1907, p. 72; Guinot, 1964, p. 89; T. Sakai, 1935, p. 71. Eriphia smithi: T. Sakai, 1976, p. 478, pl. 172, fig. 3; Miyake, 1983, p. 131, pl. 44, fig. 3; Serène, 1984, p. 311, figs. 236, 237; Dai and Yang, 1991, p. 357, pl. 47; fig. 174; Lee and Ko, 2008, p. 19, figs. 2, 5. Eriphia smithii: Terada, 1982, pp. 255 262, fig. 1 (zoeal stages).

: : 13 :... 1F, 2F, 1M 2M 2L 3M 4M. 1L, 3L, 4L 5L. 2L 2L. 1P H. 1/2 V. 5 6. 2. 3........ 1 3. 4. 7. 1 3 6. 1... 2.. CL 25 mm, CW 33 mm. I: 1/2,. 1 2 4. 2. 1 3. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2 3 5. 1, 1 1. CL 0.64 mm, RDL 1.34 mm. :. :,,,,,. :. : : 1 ( : 16.viii.2008, ); ( : 1.ix.2006, ). : 4. Family Hypothalassiidae Karasawa and Schweitzer, 2006....

14 II : 1 ( 1 ), 2 ( 1 ). : -. Genus Hypothalassia Gistel, 1848. 2.. : 2 ( 1 ). 2. Hypothalassia armata (De Haan, 1835) Cancer (Acanthodes) armatus De Haan, 1835, p. 52. Hypothalassia armata: Kim, 1973, p. 390, pl. 29; T. Sakai, 1976, p. 469, pl. 167; Miyake, 1983, p. 125, pl. 42, fig. 4. :... 2 4. 4 2... CL 101 mm, CW 119 mm (Kim, 1973). : 50 120 m, (T. Sakai, 1976). :,,. :,. :.

: : 15 Family Menippidae Ortmann, 1893.... : 5 ( 1 ), 13 ( 1 ). :. Genus Sphaerozius Stimpson, 1858.. 4. 2.. : 2 ( 1 ). 3. Sphaerozius nitidus Stimpson, 1858 ( 2A, B) Sphaerozius nitidus Stimpson, 1858a, p. 35; Kim, 1973, p. 388, pl. 28, fig. 151; T. Sakai, 1976, p. 471, pl. 171, fig. 1; Miyake, 1983, p. 126, pl. 42, fig. 5; Dai and Yang, 1991, p. 349, pl. 46, fig. 172. Sphaerozius nitidus: Ban, 1996, pp. 1 68, figs. 4 9 (larval stages). :.. 1/3 V. 4. 2..... 4 5.. 7 3. 1. 2. CL 9 mm, CW 12 mm. I:, 1.5

16 II. 1 2 2. 2 1/2. 1 1. 1 1, 4. 2 3+3 (6). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 4. 2 3. 1 (Ban, 1996). CL 0.41 mm, RDL 1.64 mm (Aikawa, 1933). :. :,,,,,. :,,,,. : : 1 ( : 4.vi.2011, ). : 5. Ban (1996). ( : : 19.ii.2007, ). Superfamily Pilumnoidea Samouelle, 1819,,.. 7. 1. 2 1 ( 2C). 1. Galenidae ( 2F) Pilumnidae 2 2. 1, 2, 1, 2+4 3+2+3 (8) 7. 1. 1. 2 8 2. 2 3 2 4 3. Heteropilumnus ciliatus

: : 17 Pilumnopeus makianus 4. 2 5 7 5. Harrovia japonica 6 6. 2 5 Halimede fragifer 2, 3 Pilumnus longicornis 7. 2 5 Pilumnus minutus 2, 3 Pilumnopeus granulatus 8. 2 5 Pilumnus trispinosus 2 Benthopanope indica Family Galenidae Alcock, 1898... 1.... : 4 ( 2 ), 12 ( 2 ). : -. 1.. Halimede. 2 3 Parapanope Genus Halimede De Haan, 1833.. 1. 2.

18 II. 1 2. : 4 ( 1 ). 4. Halimede fragifer (De Haan, 1835) ( 3A) Cancer (Halimede) fragifer De Haan, 1835, p. 47, pl. 13, fig. 4. Halimede fragifer: Kim, 1973, p. 373, pl. 81, fig. 104; T. Sakai, 1976, p. 386, pl. 136, fig. 2. Halimede tyche (not Herbst) Rathbun, 1910, p. 353; Dai and Yang, 1991, p. 264, pl. 32; fig. 142. Halimede dofleini Balss, 1922, p. 129, pl. 2, figs. 1, 2. Halimede fragifer: Terada, 1985, pp. 30 37, fig. 1 (zoeal stages). :.,,. 1/5 2. 4. 1. 2.. 10. 6 5.. 7. 1 90. CL 20 mm, CW 24 mm. I: 1/2,. 1 2 2. 2 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2 5. 1, 1 1. CL 0.35 mm, RDL 0.76 mm (Terada, 1982). : 20 50 m. :,,,,,. :,. : : 1 ( : 1.vii.2011, ). :. Kamita (1941) ( ), Kim (1973)

: : 19 Kamita. Terada (1982) 2. Genus Parapanope De Man, 1895.... 1... 1 2 (K. Sakai, 2004). : 6 ( 1 ). 5. Parapanope euagora De Man, 1895 Parapanope euagora De Man, 1895, p. 514; T. Sakai, 1976, p. 434, pl. 156, fig. 3; Dai and Yang, 1991, p. 302, pl. 39, fig. 158. Hoploxanthus hextii Alcock, 1898, p. 126. :.. 2. 4..... 2 3..... 1. 7.. CL 11 14.3 mm, CW 16.9 20.4 mm (Dai and Yang, 1991). : 35 m (T. Sakai, 1976). :,,,,,,,. :. :. Kim and Kim (1982) ( ).

20 II Family Pilumnidae Samouelle, 1819,,,... 7. 1 2 S. : 62 ( 9 ), 383 ( 13 ). :. 1. 2, 3 2. Echinoecus Harrovia 3. 4 Zehntneriana 4. 5 Pilumnus 5. Benthopanope 6 6. Heteropilumnus 7 7. 8 Pilumnopeus 8. Actumnus Neoactumnus Genus Echinoecus Rathbun, 1894..... : 3 ( 1 ).

: : 21 6. Echinoecus nipponicus Miyake, 1939 ( 3B) Echinoecus petiti nipponicus Miyake, 1939, pp. 86, 88, 90, figs. 1 3B. Echinoecus pentagonus: T. Sakai, 1976, p. 295, pls. 81, 100, fig. 1; Takeda, 1982, p. 136, fig. 402; Miyake, 1983, pp. 55, 213, pl. 19, fig. 3; Kim and Chang, 1985, pl. 51, fig. 4. Echinoecus nipponicus: Chia et al., 1999, p. 816, figs. 2Q, 3, 4A G; Ng et al., 2008, p. 139 (only list); Lee et al., 2011, p. 191, figs. 1, 2. :. 1/3... 3..... 3 4.... 1 3 4. 4, 5. 1/3. CL 7 mm, CW 8 mm. : 15 m (Lee et al., 2011). :,. :. : : 1 ( : 17.x.2010, ). :. Kim and Chang (1985) (E. pentagonus) (E. nipponicus). Genus Harrovia Adams and White, 1849... 1. 2... 1 2. : 7 ( 2 ).

22 II 7. Harrovia elegans De Man, 1887 Harrovia elegans De Man, 1887a, p. 21, pl. 1, figs. 5, 6; Kim, 1973, p. 558, pl. 112, fig. 265; Chia and Ng, 1998, pp. 514, 523, figs. 14, 15. Not Harrovia elegans: T. Sakai, 1976, p. 299, pl. 100, fig. 166; Takeda, 1982, p. 137, fig. 404; Miyake, 1983, p. 56, pl. 19, fig. 5. =Harrovia japonica Balss, 1921. :. 4, 1, 2 3, 4... 1. 7. CL 5.5 mm, CW 6.7 mm (Kim, 1973). : (Crinoids) (Kim, 1973). :,,,,,,. :. :. 8. Harrovia japonica Balss, 1921 ( 4A, B) Harrovia japonica Balss, 1921, p. 177; Balss, 1922, p. 136, figs. 8, 9; Chai et al., 1993, p. 266 (note only); Minemizu, 2000, p. 224; K. Sakai, 2004, p. 877; Lee and Ko, 2009a, p. 125, fig. 1. Harrovia elegans: T. Sakai, 1976, pp. 298, 299, pl. 100, fig. 166; Takeda, 1982, p. 137, fig. 404; Miyake, 1983, pp. 56, 212, pl. 19, fig. 5. Harrovia japonica: Lee and Ko, 2009b, pp. 213 222, figs. 1 4 (zoeal stages). :.. 2M 5L. 4.... 3...

: : 23 1 2.. 2. 1. 7 8.. 7. 1. 90. 2. CL 8 mm, CW 13 mm. I:,. 1 3 2. 2 3. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2 5. 2 4 2 3. 1, 1 1. CL 0.53 mm, RDL 0.92 mm. : 20 m (Crinoids). :,,. :,. : : 1 ( : 2.viii.2007, ), : ( : 19.x.2008, ). : 4. Genus Actumnus Dana, 1851.,. 3. (Kim, 1973). : 7 ( 2 ).

24 II 9. Actumnus elegans De Man, 1887 Actumnus elegans De Man, 1887a, p. 47; Takeda and Miyake, 1969a, p. 97; Garth and Kim, 1983, p. 696; Kim, 1985, p. 83. :.. 2.. 4.... CL 9 mm, CW 12 mm (Kim, 1985). :. :,,,. :. :. 10. Actumnus marissinicus Takeda and Kim, 1977 Actumnus asper: Kim, 1973, p. 398, pl. 84, fig. 157. Actumnus marissinicus: Takeda and Kim, 1977, p. 135, pl. 1, fig. 1; K. Sakai, 2004. :.. 2. 2. 3.... 1, 2. 3, 4. 4. 7. 1. CL 9.5 mm, CW 13.4 mm. :. :. :. :.

: : 25 Genus Benthopanope Davie, 1989.. 2. 3.. 1 3. 1 2. : 6 ( 1 ). 11. Benthopanope indica (De Man, 1887) ( 5A, B) Heteropanope indica De Man, 1887a, p. 53, pl. 3, figs. 1, 2. Pilumnopeus indica Balss, 1933, p. 33; T. Sakai, 1976, p. 500, pl. 178, fig. 2; Dai and Yang, 1991, p. 374, pl. 50, fig. 182. Pilumnopeus indicus: Miyake, 1983, p. 138, pl. 46, fig. 5. Heteropanope (Pilumnopeus) indica: Kim, 1973, p. 400, pl. 84, fig. 158. Benthopanope indica: Davie, 1989, p. 144 (only list); K. Sakai, 2004. Benthopanope indica: Ko, 1995, pp. 280 290, figs. 2 6 (larval stages). :.., 1. 2. 3... 1 3 4. 7. 1. 2. CL 8 mm, CW 6 mm. I:.. 1 3 2. 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2. 1, 1 1. CL 0.52 mm, RDL 0.64 mm. : (Sargassum). :,,,,.

26 II :,. : : 1 ( : 6.vi.2011, ); ( : 24.vii.2010, ). : 4. Genus Hetropilumnus De Man, 1895... 2. 3.. 1 2. : 19 ( 1 ). 12. Heteropilumnus ciliatus (Stimpson, 1858) ( 6A, B) Pilumnoplax ciliata Stimpson, 1858b, p. 94. Heteropilumnus ciliatus Balss, 1933, p. 42; Kim, 1973, p. 396, pl. 30, fig. 156; T. Sakai, 1976, p. 492, pl. 176, fig. 3; Dai and Yang, 1991, p. 373, pl. 50. Heteropilumnus ciliatus: Ko and Yang, 2003, pp. 341 351, figs. 1 6 (zoeal stages). :... 2. 3... CL 9 mm, CW 13 mm. I:, 1.4. 1 4 2. 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.61 mm, RDL 1.93 mm.

: : 27 :. :,,. :,,. : : 1 ( : 15.vi.2011, ), : ( : 1.viii.2010, ). : 3. Genus Neoactumnus T. Sakai, 1965..... 7 (T. Sakai, 1965). : 2 ( 1 ). 13. Neoactumnus convexus T. Sakai, 1965 Neoactumnus convexus T. Sakai, 1965a, p. 105, fig. 4; 1976, p. 498, pl. 177, fig. 267; Miyake, 1983, p. 133, pl. 45, fig. 3; Lee et al., 2008, p. 291, fig. 1. :.. 1. 3..., 1.. CL 9 mm, CW 13 mm (Lee et al., 2008). : 25 45 m (Lee et al., 2008). :,. :. :.

28 II Genus Pilumnopeus A. Milne-Edwards, 1867... 2. 3.. 7. 1 2. : 13 ( 2 ). 14. Pilumnopeus granulatus Balss, 1933 ( 7A, B) Pilumnopeus serratifrons granulatus Balss, 1933, p. 34. Pilumnopeus granulatus: Takeda and Miyake, 1969a, p. 127, fig. 12; Ko and Takeda, 2000, p. 34, fig. 3. Pilumnopeus granulata: Ko, 1997, pp. 31 42, figs. 1 5 (larval stages). :.. 2. 3..... 7. 1. CL 10 mm, CW 15 mm. I:,. 1 3 3. 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.61 mm, RDL 0.89 mm. :. :,,,,. :.

: : 29 : : 1 ( : 15.vi.2011, ); ( : 19.vi.1993, ). : 4. 15. Pilumnopeus makianus (Rathbun, 1931) ( 8A, B) Heteropanope makiana Rathbun, 1931, p. 80, pl. 11, figs. 31, 32. Pilumnopeus makiana: T. Sakai, 1976, p. 501, pl. 178, fig. 3; Dai and Yang, 1991, p. 375, pl. 50, fig. 182. Heteropanope (Pilumnopeus) makiana: Kim, 1973, p. 402, pl. 30, fig. 159. Pilumnopeus makinaus: Ng et al., 2008, p. 411 (only list). Pilumnopeus makiana: Lee, 1993, pp. 1 53, figs. 6 9 (zoeal stages). :.,. 2. 3... 7 1. CL 9 mm, CW 14 mm. I:,. 1 4 1. 2. 2. 1 1, 2+4. 2 3+2+4 (8). 1, 2+2+3+3 (10 11) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.61 mm, RDL 1.13 mm. :. :,,,. :,,,,. : : 1 ( : 1.vii.2011, ), : ( : 5.vii.2006, ). : 4. 2 2 Lee (1993) 2+5 (7) 1, 1, 5 3+2+3 (8) 1, 1, 6.

30 II Genus Pilumnus Leach, 1815.. 2... 7. 1. 2 S. : 136 ( 3 ). 16. Pilumnus longicornis Hilgendorf, 1878 Pilumnus longicornis Hilgendorf, 1879, p. 794, pl. 1, figs. 8, 9; Balss, 1933, p. 15; Kim, 1973, p. 393, pl. 84, fig. 154; T. Sakai, 1976, p. 486, pl. 175, fig. 1; Miyake, 1983, p. 138, pl. 46, fig. 4. Pilumnus longicornis: Clark and Paula, 2003, pp. 323 378, figs. 33 36 (zoea I). :.. 2. 3.... 1 3 3 5. 7. CL 13 mm, CW 18 mm (Kim, 1973). I: 2,. 1 4 2. 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. RDL 1.21 mm (Clark and Paula, 2003). : 10 20 m. :,,,,,,,,,,,,,. :. : Clark and Paula (2003) 1.

: : 31 17. Pilumnus minutus De Haan, 1835 ( 9A, B) Pilumnus minutus De Haan, 1835, p. 50, pl. 3, fig. 2; A. Milne-Edwards, 1872, p. 250; Kim, 1973, p. 395, pl. 83, fig. 155; T. Sakai, 1976, p. 487, pl. 174, fig. 260; Miyake, 1983, p. 137; Dai and Yang, 1991, p. 367, pl. 49, fig. 178. Pilumnus hirsutus: Stimpson, 1858a, p. 37; Balss, 1933, p. 20. Pilumnus minutus: Ko, 1994a, pp. 145 155, figs. 1 4 (zoeal stages); 1997, pp. 31 42, fig. 6 (megalopal stage). :... 2. 3..... 7. 1. CL 8 mm, CW 11 mm. I:,. 1 3 3. 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2 5. 1, 1 1. CL 0.45 mm, RDL 0.71 mm. : 50 m (Kim, 1973). :,,,,,,,,,,,. :,,,. : : 1 ( : 4.vi.2011, ); : ( : 1.viii.2010, ). : 4. 18. Pilumnus trispinosus (T. Sakai, 1965) ( 10A, B) Parapilumnus trispinosus T. Sakai, 1965b, p. 160, pl. 71, fig. 5; Takeda and Miyake, 1969a, p. 139, fig. 15; Sakai, 1976, p. 502, fig. 268; Kim and Chang, 1985, p. 55; Dai and Yang, 1991, p. 376, fig. 182.

32 II Pilumnus trispinosus: Ng et al., 2008, p. 142 (only list). Parapilumnus trispinosus: Ko, 1994b, pp. 331 342, figs. 1 5 (larval stages). :... 2. 3...... 7. 1. CL 7 mm, CW 11 mm. I:.. 1 3 3. 2. 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2 5. 1, 1 1. CL 0.51 mm, RDL 0.77 mm. : (Kim, 1973). :,,,,,. :. : : 1 ( : 14.vi.2011, ); ( : 19.vi.1993, ). : 4. Genus Zehntneriana Ng and Takeda, 2010.. 2. 4... 7. 1 (Takeda, 1972). : 4 ( 1 ).

: : 33 19. Zehntneriana amakusae (Takeda and Miyake, 1969) ( 11A) Lithocheira amakusae Takeda and Miyake, 1969c, p. 10, fig. 1. Zehntneria amakusae: Yamaguchi et al., 1987, p. 57, pl. 10, fig. 4; Ng et al., 2008, p. 144 (only list); K. Sakai, 2004. Zehntneriana amakusae: Ng and Takeda, 2010, p. 49; Lee et al., 2011, p. 193, figs. 3, 4. :. 1.4.,.. 3.. 3. 2..,... 3..... 1 6 1.8.. 7. 1. 1 1/3. 60. 2 1. CL 6 mm, CW 8 mm (Kim, 1973). : 15 m. :,. :. : : 1 ( : 13.viii.2010, ). :. Superfamily Xanthoidea MacLeay, 1838 (Xanthidae MacLeay, 1838).

34 II Family Xanthidae MacLeay, 1838,,.. 4. 5 3 5. 1 2 1 ( 2D). : 120 ( 21 ), 637 ( 25 ). :. 1. Palapedia 2 2.. Lybia. 3 3. 4 5 4. 4 Banareia Calvactaea 5. 1.7 6 1.7 7 6. Neoliomera Liomera 7. 8 9 8. Neoxanthops Cycloxanthops 9. Medaeops 10 10. 11 13 11. Atergatis 4 12 12. Atergatopsis Platypodia

: : 35 13. 14 19 14. 15 Pilodius 15. 16 Actaea 16. 2M 2 17 2M 2 Novactaea 17. 2M 2 Actaeodes 2M 2 18 18. Forestia Gaillardiellus 19. Leptodius 20 20. Nanocassiope 21 21. 1/2 Microcassiope 1/2 Macromedaeus,. 2 1/3 1 3. 1, 2, 1, 2+4 3+2+3 ( 2). 2, 3. 1, 1 1. 1. 2 1/4 Medaeops granulosus 2 1/4 2 2. 2 3+2+2 Palapedia integra 2 3+2+3 3 3. 2 1, 1, 6 4 2 1, 1, 5 10 4. 2 1 Actaea semblatae 2 2 3 5 5. 2 Gaillardiellus orientalis 2 6 6. 2 Novactaea pulchella 2 7 7. 2 2 Pilodius nigrocrinitus 2 3 8

36 II 8. 2 3 4 Atergatis reticulatus 2 20 9 9. 2 Actaeodes hirsutissimus 2 Atetgatis flordius 10. 2 Macromedaeus distinguendus 2 11 11. 2 1 12 2 2 13 12. 1.5 Nanocassiope granulipes 1.5 Leptodius exaratus 13. Calvactaea tumida Cycloxanthops truncatus Genus Actaea De Haan, 1833. 1.7..... : 32 ( 2 ). 20. Actaea polyacantha (Heller, 1861) Chlorodius polyacantha Heller, 1861, p. 339, pl. 3, fig. 21. Actaea polyacantha: Ortmann, 1893, p. 455; Guinot, 1976, p. 236, fig. 40B, b, pl. 13, fig. 6; T. Sakai, 1976, p. 443, fig. 234; Miyake, 1983, p. 116, pl. 39, fig. 4; Serène, 1984, p. 114, pl. 14E, fig. 67; Dai and Yang, 1991, p. 309, fig. 160; Lee et al., 2010, p. 87, fig. 1. :.. 2M 3M. 2 V. 6.

: : 37. 3 4. 7. CL 11 mm, CW 16 mm (Lee et al., 2010). : (Lee et al., 2010). :,,,,,,. :. :. 21. Actaea semblatae Guinot, 1976 ( 12A, B) Cancer (Actaea) granulatus (not Linnaeus, 1758) De Haan, 1835, p. 47, pl. D. Actaea granulata carcharias T. Sakai, 1934, p. 309. Actaea savignyi (not H. Milne-Edwards, 1834): T. Sakai, 1935, p. 70; 1976, p. 442, pl. 158, fig. 2; Kim, 1973, p. 383, pl. 83, fig. 147. Actaea semblatae Guinot, 1976, p. 225, pl. 11, fig. 39c; Miyake, 1983, p. 115, pl. 39, fig. 2; Serène, 1984, pp. 109, 112; Kim and Chang, 1985, p. 54. Actaea semblatae: Ko et al., 2002, pp. 1 12, figs. 1 4 (zoeal stages). :.. 2 V..... 5 3 5. CL 19 mm, CW 26 mm. I: 2,. 1 4 2. 2.. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 2 1. CL 0.70 mm, RDL 3.49 mm. :,,. : 10 m.

38 II :,,. : : 1 ( : 16.vi.2009, ), : ( : 1.viii.2010, ). : 2. Genus Actaeodes Dana, 1852. 1.7.. 2M 2.. 4... : 6 ( 1 ). 22. Actaeodes hirsutissimus (Rüppell, 1830) Xantho hirsutissimus Rüppell, 1830, p. 26, pls. 5, 6, fig. 6. Cancer (Actaea) hirsutissima De Haan, 1833, p. 18. Actaeodes hirsutissimus: Guinot, 1967a, p. 561; 1976, p. 245, fig. 38E, pl. 15; T. Sakai, 1976, p. 448, pl. 159, fig. 3; Serène, 1984, pp. 133 135, fig. 79, pl. 18B; Dai and Yang, 1991, p. 313, pl. 40, fig. 161; Lee et al., 2010, p. 90, fig. 2. Actaeodes hirsutissimus: Clark and Al-Aidaroos, 1994, pp. 207 214, figs. 1a, b, d, e, h, 2a d, 3a d (zoea I). : 1.5 1.6... 2M 2. 2 V. 4.... 3 5. 1. CL 9 mm, CW 14 mm (Lee et al., 2010). I:. 2.. 1 3 2. 2

: : 39 20. 1/8 1. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. (Clark and Al-Aidaroos, 1996). : (Lee et al., 2010). :,,,,,. :. : 1 CL RDL. Genus Forestia Guinot, 1976. 1.7.. 2M 2.... : 4 ( 1 ). 23. Forestia depressa (White, 1848) ( 11B) Xantho depressa White, 1848, p. 2. Actaea depressa: Odhner, 1925, p. 38, pl. 2, fig. 19; Guinot, 1967a, p. 261; T. Sakai, 1976, p. 446, fig. 237. Forestia depressa: Guinot, 1976, p. 262, figs. 42B, 44A, 45; Miyake, 1983, p. 115; Serène, 1984, p. 106, pl. 15, fig. 64; Ko and Takeda, 1999, p. 76, fig. 1. : 1.4.., 2M 2. 2 V. 4... 7. CL 24 mm, CW 35 mm.

40 II : 13 m. :,,,,,,,. :. : : 1 ( : 12.viii.2010, ). :. Genus Gaillardiellus Guinot, 1976. 1.7... 2M 2. 3 4.... : 5 ( 1 ). 24. Gaillardiellus orientalis (Odhner, 1925) ( 13A, B) Actaea ruppelli var. orientalis Odhner, 1925, p. 46, pl. 3, fig. 7. Actaea rüppelli orientalis: T. Sakai, 1935a, p. 70. Paractaea ruppelli orientalis: Kim, 1960, p. 343; T. Sakai, 1976, p. 452, pl. 159, fig. 2. Actaea rueppelli orientalis: Kim, 1973, p. 385, pl. 27, fig. 112. Gaillardiellus orientalis: Guinot, 1976, p. 255, fig. 43B, b, pl. 16, fig. 2; Miyake, 1983, p. 116, pl. 39, fig. 6; Serène, 1984, p. 118 (keys); Kim and Chang, 1985, p. 54. Paractaea orientalis: Dai and Yang, 1991, p. 315, pl. 40. Gaillardiellus orientalis: Ko, 1999, pp. 51 65, figs. 1 7 (larval stages). : 1.3.. 2M 2. 2 V. 4 2..... 3 5.

: : 41 1. CL 18 mm, CW 25 mm. I: 2,. 1 4 2. 2. 1/10 3. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 2, 2. CL 0.59 mm, RDL 2.67 mm. : 15 m. :,,. :,,,. : : 1 ( : 6.vi.2011, ), : ( : 1.viii.2010, ). : 4. Genus Novactaea Guinot, 1976. 1.7... 2M 2..... : 4 ( 1 ). 25. Novactaea pulchella (A. Milne-Edwards, 1865) ( 14A) Actaea pulchella A. Milne-Edwards, 1865, p. 273, pl. 18, fig. 5; Guinot, 1967a, p. 260; T. Sakai, 1976, p. 445, fig. 236; Dai and Yang, 1991, p. 307, pl. 39, fig. 159. Novactaea pulchella: Guinot, 1976, p. 269, pl. 18, fig. 6; Miyake, 1983, p. 116, pl. 39, fig. 5; Serène, 1984, pp. 104 (key), 105, pl. 14A; Ko and Takeda, 2000, p. 32, fig. 1. Novactaea pulchella: Ko, 2006, pp. 7 14, figs. 1 3 (larval stages).

42 II : 1.6... 2M 2. 2.... 3 4... 7. CL 9 mm, CW 15 mm. I:,. 1 4 1.. 2. 1/4 2.. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3 2 5. 1, 1 1. CL 0.92 mm, RDL 3.09 mm. : 15 m. :,,,,,,. :. : : 1 ( : 15.vi.2011, ); ( : 17.viii.1998, ). : 2. 2, 3 2 5 2 (Ko, 2006). Genus Pilodius Dana, 1851. 1.7... 4.... : 17 ( 2 ).

: : 43 26. Pilodius miersi (Ward, 1936) ( 14B) Chlorodopsis miersi Ward, 1936, p. 4, pl. 2, figs. 1 3. Pilodius granulatus: T. Sakai, 1976, p. 460, pl. 164, fig. 3; Serène, 1984, p. 239 (key); Dai and Yang, 1991, p. 329, pl. 43, fig. 165. Pilodius miersi: Clark and Galil, 1993, p. 1136, figs. 7A G, 34A, 41D; Lee and Ko, 2011, p. 187, figs. 5, 6. : 1.7.. 2M 2. 2. 4 1 4... 2... 3 5. 1. 2. CL 8 mm, CW 13 mm. : 25 m. :,,,,,,. :. : : 1 ( : 16.vi.2009, ). :. 27. Pilodius nigrocrinitus Stimpson, 1859 ( 15A, B) Pilodius nigrocrinitus Stimpson, 1858a, p. 34; T. Sakai, 1976, p. 461, pl. 164, fig. 2; Miyake, 1983, p. 121, pl. 41, fig. 4; Serène, 1984, p. 239 (key); Dai and Yang, 1991, p. 330, pl. 43, fig. 166; Clark and Galil, 1993, p. 1139, figs. 9A G, 35A, 42B C; Ko and Takeda, 1999, p. 77, fig. 2. Chlorodopsis melanochirus: A. Milne Edwards, 1873, p. 228, pl. 8, fig. 5. Chlorodopsis melanochira: De Man, 1887b, p. 281; Forest and Guinot, 1961, p. 89. Chlorodopsis nigrocrinita: Alcock, 1898, p. 168. Pilodius nigrocrinitus: Terada, 1981, pp. 23 28, fig. 1 (zoeal stages).

44 II : 1.5.. 2M 2. 2. 4 2 4.. 2..... 3 5. 1. CL 8 mm, CW 12 mm. I:,. 1 3 1. 2. 1/10 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.56 mm, RDL 1.42 mm. :. :,,,,,,,,,,,,,,. :. : : 1 ( : 15.vi.2011, ); ( : 12.viii.2010, ). : 3. Genus Medaeops Guinot, 1967. 1.7.. 4M. 4.... : 6 ( 1 ).

: : 45 28. Medaeops granulosus (Haswell, 1882) ( 16A) Leptodius granulosus Haswell, 1882, p. 61. Medaeus granulosus: T. Sakai, 1935, p. 70; Kamita, 1941, p. 118, fig. 62; Kim, 1970, p. 14. Medaeops granulosus: Guinot, 1967b, p. 366; Kim, 1973, p. 382, pl. 82, fig. 110; T. Sakai, 1976, p. 425, pl. 153, fig. 3; Miyake, 1983, p. 109, pl. 37, fig. 3; Serène, 1984, p. 91 (key); Dai and Yang, 1991, p. 295, pl. 37, fig. 155. Medaeops granulosus: Terada, 1990, pp. 23 47, figs. 1, 2 (zoeal stages). : 1.5.. 4M. 2. 4...... 3 5. 1. 2. CL 13 mm, CW 20 mm. I:,. 1 2 1. 2 1/2. 1/3 3. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 5. 2, 3. 1 1. CL 0.60 mm, RDL 2.04 mm. :. :,,,,,,. :,,,. : : 1 ( : 6.vi.2011, ); ( : 6.vi.2011, ). : 4.

46 II Genus Palapedia Ng, 1993..... : 15 ( 1 ). 29. Palapedia integra (De Haan, 1835) ( 16B) Cancer (Xantho) integra De Haan, 1835, p. 66, pl. 18, fig. 6. Kraussia integra: T. Sakai, 1976, p. 308, fig. 172; Dai and Yang, 1991, p. 191, pl. 23, fig. 103. Palapedia integra: Ng, 1993, p. 141; Ko and Takeda, 1999, p. 78, fig. 3. Palapedia integra: Ko et al., 2004, pp. 637 651, figs. 2 9 (zoeal stages). :,... 4 2 2...... 3 5 6. 1. 2. CL 14 mm, CW 16 mm. I:,. 1 4 1. 2. 1/10 2. 1 1, 2+4. 2 3+2+2 (7). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.47 mm, RDL 1.58 mm. : 10 m. :,,,,. :. : : 1 ( : 23.v.2009, ); ( : 7.vii.2002, ).

: : 47 : 2 3+2+2 (7). 4. Genus Liomera Dana, 1851. 1.7.. 4.. : 29 ( 1 ). 30. Liomera margaritata (A. Milne-Edwards, 1873) ( 17A) Carpilodes margaritatus A. Milne-Edwards, 1873, p. 182, pl. 5, fig. 2. Liomera margaritata: T. Sakai, 1965b, p. 144, pl. 72, fig. 3; 1976, p. 396, fig. 211; Guinot, 1967a, p. 266; Dai and Yang, 1991, p. 271, pl. 34, fig. 145; Lee and Ko, 2011, p. 185, figs. 1, 2. Liomera (Liomera) margaritata: Serène, 1984, p. 63, fig. 23, pl. 7A. : 1.7... 1M 2M 2M 2. 2. 4..... 3 5 6. 1. 2. CL 14 mm, CW 24 mm. : 25 m. :,,,,,,,,,,. :. : : 1 ( : 16.vi.2009, ). :.

48 II Genus Neoliomera Odhner, 1925. 1.7.... : 16 ( 1 ). 31. Neoliomera insularis (Adams and White, 1849) ( 17B) Atergatis insularis Adams and White, 1848, p. 38, pl. 8, fig. 2. Neoliomera insularis: T. Sakai, 1936, p.159, pl. 48 fig. 4; 1976, p. 398, pl. 141, fig. 2; Kim, 1973, p. 374, pl. 82, fig. 105; Serène, 1984, p. 67 (key). : 1.7.. 2........ 7. CL 10 mm, CW 17 mm. : 15 m. :,,,,,,,,. :. : : 1 ( : 13.viii.2010, ). :. Genus Lybia H. Milne-Edwards, 1834.... 1, 2.

: : 49 : 10 ( 1 ). 32. Lybia caestifera (Alcock, 1898) Melia caesifer Alcock, 1898, p. 231; 1899, pl. 38, fig. 4. Lybia caestifera: T. Sakai, 1976, p. 504, pl. 180, fig. 2; Guinot, 1976, p. 75, pl. 19, fig. 5; Miyake, 1983, p. 128, pl. 43, fig. 6; Serène, 1984, p. 25 (list); Dai and Yang, 1991, p. 379, pl. 51; Lee et al., 2008, p. 294, fig. 2. :. 1F, 2M 4L.. 3.... CL 4 mm, CW 4 mm (Lee et al., 2008). :. :,,,,. :. :. Genus Cycloxanthops Rathbun, 1897. 1.7....... : 6 ( 1 ).

50 II 33. Cycloxanthops truncatus (De Haan, 1837) ( 18A, B) Cancer (Xantho) truncatus De Haan, 1837, p. 66, pl. 18, fig. 4. Xantho truncata: T. Sakai, 1935, p. 71. Cycloxanthops truncatus: T. Sakai, 1976, p. 435, fig. 229; Miyake, 1983, p. 108, pl. 36, fig. 5; Kim and Chang, 1985, p. 54. Cycloxanthops truncatus: Terada, 1980, pp. 138 148, figs. 1b, 2b, 3b (larval stages). : 1.5.. 2. 5....... 3 5 6. 1 90. 2. CL 15 mm, CW 22 mm. I: 0.7, 1. 1 2 1. 2. 1/8 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 5. 2, 3. 1, 1 1. CL 0.53 mm, RDL 1.56 mm. :. :,. :. : : 1 ( : 16.vi.2011, ); ( : 13.viii.2010, ). : 4. Genus Leptodius A. Milne-Edwards, 1863. 1.7.

: : 51. 4.... : 12 ( 1 ). 34. Leptodius exaratus (H. Milne-Edwards, 1834) ( 19A, B) Chlorodius exaratus H. Milne-Edwards, 1834, p. 402. Leptodius exaratus: Balss, 1922, p. 127; T. Sakai, 1935, p. 71; 1976, p. 423, pl. 153, fig. 1; Kim, 1973, p. 380, pl. 82, fig. 109; Miyake, 1983, p. 109, pl. 37, fig. 2; Serène, 1984, p. 183, pl. 26A, fig. 106; Kim and Chang, 1985, p. 53; Dai and Yang, 1991, p. 292, pl. 37, fig. 154. Leptodius exaratus: Terada, 1980, pp. 138 148, figs. 1a, 2a, 3a (zoeal stages). : 1.5.. 2. 4...... 3 5. 1 45. 2. CL 20 mm, CW 30 mm. I: 0.7, 1. 1 2 2. 2. 1/10 1. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+ 1+1+1 (4) 1, 1, 5. 2, 3. 1, 1 1. CL 0.50 mm, RDL 1.16 mm. :. :,,,,,,,,,,,,,,,,. :.

52 II : : 1 ( : 15.vi.2011, ); ( : 24.vii.2010, ). : 4. Genus Macromedaeus Ward, 1942. 1.7.. 4... 1/2... : 6 ( 1 ). 35. Macromedaeus distinguendus (De Haan, 1835) ( 20A, B) Cancer (Xantho) distinguendus De Haan, 1835, p. 48, pl. 13, fig. 7. Xantho distinguendus: T. Sakai, 1935, p. 71. Leptodius distinguendus: T. Sakai, 1965b, p. 141, pl. 70, fig. 3. Macromedaeus distinguendus: Kim, 1973, p. 379, pl. 27, fig. 108; T. Sakai, 1976, p. 419, fig. 221, pl. 153, fig. 2; Miyake, 1983, p. 110, pl. 37, fig. 5; Serène, 1984, p. 177 (key); Dai and Yang, 1991, p. 286, pl. 36, fig. 151. Macromedaeus distinguendus: Lee, 1993, pp. 1 53, figs. 1 5 (larval stages). : 1.5.., 6L. 2. 4... 1/2..... 3 5. 1. 2. CL 17 mm, CW 25 mm.

: : 53 I: 1.3,. 1 3 2. 2. 1/10 1. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 5. 2, 3. 1, 1 1. CL 0.55 mm, RDL 1.79 mm. :. :,,,,,. :,,,,,,. : : 1 ( : 1.vii.2011, ); ( : 30.vi.2011, ). : 4. Genus Microcassiope Guinot, 1967.. 1.7... 4... 1/2... : 5 ( 1 ). 36. Microcassiope orientalis Takeda and Miyake, 1969 ( 21A, B) Microcassiope orientalis Takeda and Miyake, 1969b, p. 201, figs. 2, 3. Macromedaeus orientalis: Takeda, 1977, p. 84; Lee and Ko, 2008, p. 17, figs. 1, 4. Macromedaeus orientalis (not Microcassiope orientalis): Ko, 2002, pp. 89 93, figs. 1, 2 [zoea I=Macromedaeus distinguendus (De Haan, 1835)]. : 1.5... 1F

54 II 2F. 2M. 1L 5L. 6L. 3M 4M. 2. 4.... 1/2..... 3 5. 1. 2. CL 9 mm, CW 14 mm. : 20 m. :,. :. : : 1 ( : 15.vi.2011, ). :. Genus Nanocassiope Guinot, 1967.. 1.7... 4..... : 6 ( 1 ). 37. Nanocassiope granulipes (T. Sakai, 1939) ( 22A) Heteropanope granulipes T. Sakai, 1939, p. 546, fig. 59. Micropanope granulipes: T. Sakai, 1965b, p. 139, pl. 70, fig. 2. Nanocassiope granulipes: Takeda and Miyake, 1968, p. 552; Kim and Chang, 1985, p. 54. Nanocassiope granulipes: Ko and Clark, 2002, pp. 1463 1488, figs. 1 13 (zoeal stages).

: : 55 : 1.6.. 3M.. 4...... 3 5. 1. 2. CL 7 mm, CW 11 mm. I: 1.3, 1.6. 1 4 1. 2. 1/9 1. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 5. 2, 3. 1, 1 1. CL 0.46 mm, RDL 1.50 mm. : 20 m. :,. :. : : 1 ( : 13.viii.2010, ); ( : 13.vii.1999, ). : 4. Genus Neoxanthops Guinot, 1968. 1.7....... : 3 ( 1 ).

56 II 38. Neoxanthops lineatus (A. Milne-Edwards, 1867) ( 22B) Cycloxanthus lineatus A. Milne-Edwards, 1867, p. 269. Cycloxanthops lineatus: T. Sakai, 1939, p. 454, pl. 55, fig. 2. Neoxanthops lineatus: Guinot, 1968, p. 700, fig. 16; T. Sakai, 1976, p. 436, pl. 157, fig. 1; Serène, 1984, pp. 211 (key), 212, pl. 29C, fig. 127; Lee and Ko, 2011, p. 185, figs. 3, 4. : 1.6.. 2. 3 4..... 7. CL 12 mm, CW 19 mm. : 25 m. :,,,,,,,,. :. : : 1 ( : 16.vi.2009, ). :. Genus Banareia A. Milne-Edwards, 1869.. 4... : 16 ( 1 ). 39. Banareia subglobosa (Stimpson, 1858) Actaea subglobosa Stimpson, 1858a, p. 33; Kim, 1973, p. 386, pl. 28, fig. 113.

: : 57 Actaea (Banareia) subglobosa: T. Sakai, 1939, p. 494, pl. 94, fig. 9. Banareia subglobosa: T. Sakai, 1976, p. 517, pl. 187, fig. 2; Guinot, 1976, p. 146, figs. 24F, 33A B, pl. 7, figs. 8 10; Miyake, 1983, p. 143, pl. 48, fig. 3; Serène, 1984, pp. 39 (key), 40 (key); Kim and Chang, 1985, p. 54; Dai and Yang, 1991, p. 390, pl. 53, fig. 189. : 1.3.. 2. 3 4... 7. CL 29 mm, CW 40 mm. : 10 35 m (K. Sakai, 2004). :,,,,,. :. : : 1 ( : 22.ix.2009, ). :. Genus Calvactaea Ward, 1933..... : 1 ( 1 ). 40. Calvactaea tumida Ward, 1933 Calvactaea tumida Ward, 1933, p. 384, pl. 23, fig. 9; T. Sakai, 1976, p. 520, pl. 187, fig. 3; Guinot, 1976, p. 185, figs. 25E, 26C, 27E, 34F G, pl. 7, figs. 4, 5; Dai and Yang, 1991, p. 391, pl. 53; Ko and Takeda, 2000, p. 33, fig. 2. Calvactaea tumida: Terada, 1988, pp. 43 50, fig. 1 (zoeal stages). : 1.2.

58 II.... 3.. 7. 1. CL 8 mm, CW 9 mm. I: 1.4, 1.7. 1 3 1. 2. 1/10 2. 1 1, 2+4. 2 3+2+3 (8). 1, 2+2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 5. 2, 3. 1, 1 1. CL 0.58 mm, RDL 2.21 mm (Terada, 1988). : 10 30 m (Alcyonarians). :,,,,,. :. : : 1 ( : 15.vii.1999, ). : Terada (1988) 3. Terada 3 4. Genus Atergatis De Haan, 1833. 1.7... 4... : 16 ( 2 ).

: : 59 41. Atergatis floridus (Linnaeus, 1767) ( 23A) Cancer floridus Linnaeus, 1767, p. 1041. Cancer (Atergatis) floridus De Haan, 1835, p. 46. Atergatis floridus Dana, 1852, p. 159; T. Sakai, 1936, p. 147, pl. 45, fig. 1; 1976, p. 409, pl. 148, fig. 2; Guinot, 1967a, p. 261; Kim, 1973, p. 375, pl. 25, fig. 106; Miyake, 1983, p. 101, pl. 34, fig. 1; Serène, 1984, pp. 145 (key), 146 (key), 148, pl. 21D, fig. 87; Dai and Yang, 1991, p. 282, pl. 35, fig. 150. Atergatis floridus: Tanaka and Konishi, 2001, pp. 21 42, figs. 1 12; 2009, pp. 55 63, figs. 1 4 (larval stages). : 1.4.. 2....... 3 5. 1. 2. CL 39 mm, CW 57 mm. I:,. 1 4 1. 2 20. 1/8 3. 1 1, 2+4. 2 3+2+3 (8). 1, 2+ 2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.55 mm, RDL 1.40 mm (Tanaka and Konishi, 2001; 2009). : 10 m. :,,,,,,,,,,,,,,,. :. : : 1 ( : 12.viii.2010, ). : 4.

60 II 42. Atergatis reticulatus (De Haan, 1835) ( 23B) Cancer (Atergatis) reticulatus De Haan, 1835, p. 47. Atergatis reticulatus: T. Sakai, 1936, p. 148, pl. 43, fig. 1; 1976, p. 411, pl. 147, fig. 1; Kim, 1973, p. 377, pl. 26, fig. 107; Miyake, 1983, p. 101, pl. 34, fig. 4; Serène, 1984, pp. 145 (key), 146 (key); Dai and Yang, 1991, p. 283, pl. 36, fig. 150. Atergatis reticulatus: Terada, 1980, pp. 138 148, figs. 1d, 2d, 3d (zoeal stages). : 1.6.. 2....... 3 5. 1.. CL 40 mm, CW 65 mm. I:,. 1 3 1. 2 3 4. 1/10 3. 1 1, 2+4. 2 3+2+3 (8). 1, 2+ 2+3+3 (10) 3, 2, 1, 2, 5. 2, 1+1+1+1 (4) 1, 1, 6. 2, 3. 1, 1 1. CL 0.58 mm, RDL 1.58 mm (Terada, 1980). : 20 m. :,,,. :,. : : 1 ( : 15.vii.2003, ). : 4. Genus Atergatopsis A. Milne-Edwards, 1862. 1.7.

: : 61.. 4... : 10 ( 1 ). 43. Atergatopsis germaini A. Milne-Edwards, 1865 Atergatopsis germaini A. Milne-Edwards, 1865, p. 257, pl. 11, figs. 1, 1a b; T. Sakai, 1976, p. 413, fig. 217; Miyake, 1983, p. 103, pl. 35, fig. 2; Serène, 1984, pp. 141 (key), 142 (key); Lee et al., 2008, p. 296, fig. 3. :... 3. 1/3 2.... 1 4. CL 9 mm, CW 15 mm (Lee et al., 2008). : (Lee et al., 2008). :,,,,,. :. :. Genus Platypodia Bell, 1835. 1.7... 4... : 10 ( 1 ).

62 II 44. Platypodia tomentosa (De Man, 1902) Lophactaea tomentosa De Man, 1902, p. 585. Platypodia tomentosa: T. Sakai, 1983, p. 15, pl. 2, fig. 4, pl. 7A; Serène, 1984, pp. 157 (key), 158 (key); Lee et al., 2008, p. 296, fig. 4. :. 1M 2M. 1F 2F 1M 2M 2. 4.... CL 6 mm, CW 8 mm (Lee et al., 2008). : (Lee et al., 2008). :,,. :. :.

63 Adams, A. and A. White, 1848. Crustacea. Part 1. In: A. Adams (ed.), The zoology of the voyage of H.M.S. Samarang, under the command of Captain Sir Edward Belcher, during the years 1843 1846, Benham and Reeve, London, pp. 1 32, pls. 1 6. Aikawa, H., 1933. On larval forms of some Brachyura, paper II; a note on indeterminable zoeas. Rec. Oceanogr. Wks. Japan, 5: 124 254. Alcock, A.W., 1898. Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part 1. The Family Xanthidae. J. Asiatic Soc. Bengal, Calcutta, 67, part 2(1): 67 233. Alcock, A.W., 1899. Crustacea. Part VII. In: Illustrations of the Zoology of the Royal Indian Marine Survey Ship Investigator, under the command of Commander T.H. Heming, R.N. Fishes, Calcutta, pls. 36 45. Balss, H., 1921. Diagnosen neuer Decapoden aus den Sammlungen der Deutschen Tiefsee-Expedition und der Japanischen Ausbeute Dofleins und Haberers. Zool. Anz., 52(6/7): 175 178. Balss, H., 1922. Östasiatische Decapoden. IV. Die Brachyrhynchen (Cancridea). Archiv für Naturgeschichte, 88A(11): 94 166, figs. 1 2, pls. 1 2. Balss, H., 1933. Beiträge zur Kenntnis der Gattung Pilumnus (Crustacea, Dekapoda) und verwandter Gattungen. Capita Zoologica, 4(3): 1 47, figs. 1 7, pls. 1 7. Ban, G.H., 1996. The complete larval development of Actaea semblatae (Guinot, 1976) and Sphaerozius nitidus (Stimpson, 1858) reared in the laboratory. Master thesis, Pusan Nat. Univ., pp. 1 68. Chia, D.G.B., P. Castro and P.K.L. Ng, 1999. Revision of the genus Echinoecus (Decapoda: Brachyura: Eumedonidae), crabs symbiotic with sea urchins. J. Crust. Biol., 19: 809 824. Chia, D.G.B. and P.K.L. Ng, 1998. A revision of Ceratocarcinus White, 1847, and Harrovia Adams & White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bull. Zool., 46(2): 493 563, figs. 1 33. Chia, D.G.B., P.K.L. Ng and D. Vandenspiegel, 1993. The identities of two crinoid symbionts, Harrovia albolineata Adams and White, 1849, and H. longipes Lanchester,1990 (Decapoda, Brachyura, Eumedonidae). Crustaceana, 64: 259 280. Clark, P.F. and A.M. Al-Aidaroos, 1994. The first zoeas of Actaeodes hirsutissimus (Rüppell, 1830) and A. tomentosus (H. Milne Edwards, 1834) (Crustacea: Decapoda: Brachyura: Xanthidae: Actaeinae). J. King Abdulaziz Univ., Mar. Sci. Special issue: Symp. on Red Sea Mar. Environ., Jeddah, 1994. 7: 207 214. Clark, P.F., D.K. Calazans and G.W. Pohle, 1998. Accuracy and standardization of brachyuran larval descriptions. Invert. Reprod. Develop., 33: 127 144. Clark, P.F. and B.S. Galil, 1993. A revision of the xanthid genus Pilodius Dana, 1851 (Crustacea: Brachyura: Xanthoidea). J. Nat. Hist., 27: 1119 1206, figs. 1 44. Clark, P.F. and J. Paula, 2003. Descriptions of ten xanthoidean (Crustacea: Decapoda: Brachyura) first stage zoeas from Inhaca Island, Mozambique. Raffles Bull. Zool., 2003 51: 323 378. Dai, A.Y. and S.L. Yang, 1991. Crabs of the China Seas. Springer-Verlag, Verlin, Heidelberg, New York, Tokyo, pp. 1 608, pls. 1 74. Dana, J.D., 1852. Crustacea. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U.S.N., 13(pt. 1): i viii, 1 685, pl. 8. Davie, P.J.F., 1989. A re-appraisal of Heteropanope Stimpson, and Pilumnopeus A. Milne Edwards (Crustacea: Decapoda: Pilumnidae) with descriptions of new species and new genera. Mem. Qld. Mus., 27(2): 129 156. De Haan, W., 1833 1850. Crustacea, in P.F. v. Siebold. Fauna Japonica, sive Descriptio animalium, quae in

64 II itinere per Japoniam, jussu et auspiciis superiorum, qui summum in India Batava imperium tenent, suscepto, annis 1823 1830 collegit, notis, observationibus et adumbrationibus illustravit: 1 243. Lugduni- Batavorum, Leiden. De Man, J.G., 1887a. Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F.R.S. Superintendent of the Museum. Part I. J. Linn. Soc. London., (Zool.), 22(136): 1 64, pls. 1 3. De Man, J.G., 1887b. Bericht über die von Herrn. Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv für Naturgeschichte, Berlin, 53(1): 215 288, pls. 7 10. De Man, J.G., 1895. Bericht über die von Herrn Schiffscapitün Storm zu Atjeh, an den westlichen Küsten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. (In 6 parts). Zool. Jahr., Abteilung für Systematik, Geographie und Biologie der Thiere. 8: 485 609, figs. 1 15, pls. 12 14. De Man, J.G., 1902. Die von Herrn Professor Kükenthal im indischen Archipel gesammelten Dekapoden und Stomatopoden. In: W. Kükenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo, in Aufträge der Senckenberg. Naturforsch. Gesellschaft ausgeführt von Dr. Willy Kükenthal. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25(3): 465 929, pls. 19 27. Forest, J. and D. Guinot, 1961. Crustacés Décapodes Brachyoures de Tahiti et des Tuamotu. In: Expédition française sur les récifs coralliens de la Nouvelle Calédonie. Vol. Préliminaire: i xi, 1 195, figs. 1 178, pls. 1 18, 7 maps, tables. 1 3. Garth, J.S. and H.S. Kim, 1983. Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U.S. Fish Commission Steamer Albatross in 1908 1909. J. Nat. Hist., London, 17: 663 729. Guinot, D., 1964. Crustacés Décapodes Brachyoures (Xanthidae) des campagnes de la Calypso en Mer Rouge (1952), dans le Golfe Persique et á l île Aldabra (1954). Mém. Mus. Nat. Hist. nat., Paris, (A) 32(1): i iii, 1 108, figs. 1 57, pls. 1 12. Guinot, D., 1967a. La faune carcinologique (Crustacea Brachyura) de l Océan Indien Occidental et de la Mer Rouge: Catalogue, remarques biogéographiques et bibliographiques. In: Réunion de Spécialistes C.S.A. sur les Crustacés, Zanzibar 1964. Mem. I. F. A. N., 77 (1966): 235 352, pls. 1 26. Guinot, D., 1967b. Les anciens genres Micropanope Stimpson et Medaeus Dana. In: Recherches Préliminaires sur les groupements naturels chez les Crustacés Décapodes Brachyoures. II. Bull. Mus. Nat. Hist. nat., Paris, (2) 39(2): 345 374, figs. 1 42. Guinot, D., 1968. Observations sur quelques genres de Xanthidae. In: Recherches préliminaires sur les groupements naturels chez les Crustacés, Décapodes, Brachyoures. IV. Bull. Mus. Nat. Hist. nat., Paris, (2) 39(4), (1967): 695 727, figs. 1 60. Guinot, D., 1976. Constitution de quelques groupes naturels chez les Crustacés Décapodes Brachyoures. I. La superfamille des Bellioidea Dana et trois sous-familles de Xanthidae (Polydectinae Dana, Trichiinae de Haan, Actaeinae Alcock). Mém. Mus. Nat. Hist. nat., Paris, séries A, 97: 1 308, figs. 1 47, pls. 1 19. Haswell, W.A., 1882. Catalogue of the Australian stalk- and sessile-eyed Crustacea. The Australian Museum, Sydney: i-xxiv, 1-324 + 2 p. addenda, figs. 1 8, pls. 1 4. Heller, C., 1861. Beiträge zur Crustaceen-Fauna des Roten Meeres. Part 1. Sitzungsber. Math.-nat. Cl. K. Akad. Wiss. Wien., 43(1): 297 374, pls. 1 4. Hilgendorf, F., 1879. Die von Herrn W. Peters in Moçambique gesammelten Crustaceen. Monatsber. K. Akad. Wiss., Berlin, 1878: 782 851, pls. 1 4. Kamita, T., 1941. Studies of the decapod crustaceans of Chosen. Pt. I. Crabs. Fish. Soc. Chosen, Keijo, pp. 1 289, figs. 1 147, pls. 2. Kim, H.S., 1960. The crabs and hermit-crabs from Dagelet island and Dok-do. J. Korean Cult. Res. Inst., Ewha

65 Women s Univ., 1: 341 344. Kim, H.S., 1970. A checklist of the Anomura and Brachyura (Crustacea, Decapoda) of Korea. Seoul Univ. J., Biol. Agri., (Ser. B), 21: 1 34, fig. 1, pls. 1 5. Kim, H.S., 1973. Anomura, Brachyura. Illustrated encyclopedia of fauna and flora of Korea. The Ministry of Education, Korea, 14: 1 506. Kim, H.S., 1985. Systematic studies of Crustaceans of Korea. 1. Decapods. Proc. Coll. Natur. Sci., Seoul Nat. Univ., 1: 63 94. Kim, H.S. and C.Y. Chang, 1985. The brachyuran crab of Cheju Island, Korea (Crustacea: Decapoda). Korean J. Syst. Zool., 1: 41 60. Kim, W. and H.S. Kim, 1982. Classification and geographical distribution of Korean crabs (Crustacea, Decapoda, Brachyura). Proc. Coll. Nat. Sci., Seoul Nat. Univ., 7: 133 159. Ko, H.S., 1994a. The zoeal stages of Pilumnus minutus De Haan, 1835 (Decapoda: Brachyura: Pilumnidae) in the laboratory. Korean J. Syst. Zool., 10: 145 155. Ko, H.S., 1994b. Larval development of Parapilumnus trispinosus Sakai, 1965 (Crustacea, Brachyura, Xanthidae) reared in the laboratory. Korean J. Zool., 37: 331 342. Ko, H.S., 1995. Larval development of Benthopanope indica (De Man, 1887) (Decapoda: Brachyura: Pilumnidae) in the laboratory. J. Crust. Biol., 15: 280 290. Ko, H.S., 1997. Larval development of Pilumnopeous granulata Balss, 1933 and Pilumnus minutus De Haan, 1835 (Crustacea: Brachyura: Pilumnidae), with a key to the known pilumnid zoeae. Korean J. Biol. Sci., 1: 31 42. Ko, H.S., 1999. Complete larval development of Gaillardiellus orientalis (Odhner, 1925) (Crustacea: Decapoda: Xanthidae) reared in the laboratory. Korean J. Syst. Zool., 15: 51 65. Ko, H.S., 2002. First zoeal stage of Macromedaeus orientalis (Takeda et Miyake, 1969) (Crustacea: Decapoda: Xanthidae) reared in the laboratory. Korean J. Biol. Sci., 6: 89 93. Ko, H.S., 2006. Complete larval development of Novactaea pulchella (Crustacea: Decapoda: Xanthidae). Integr. Biosci., 10: 7 14. Ko, H.S. and P.F. Clark, 2002. The zoeal development of Nanocassiope granulipes (Sakai, 1939) (Crustacea: Decapoda: Brachyura: Xanthidae) described from laboratory-reared material. J. Nat. Hist., 36: 1463 1488. Ko, H.S. and M. Takeda, 1999. New records of three xanthid crabs (Decapoda: Brachyura: Xanthidae) in Korea. Korean J. Syst. Zool., 15: 75 82. Ko, H.S. and M. Takeda, 2000. New records of three xanthoid crabs (Decapoda, Brachyura) collected from Chejudo Island in Korea. Korean J. Syst. Zool., 16: 31 37. Ko, H.S., H.S. An and S. Sulkin, 2004. Zoeal development of Palapedia integra (De Haan, 1835) (Crustacea: Decapoda: Brachyura: Xanthidae) reared in the laboratory. J. Crust. Biol., 24: 637 651. Ko, H.S., H.J. Yang and K.H. Ban, 2002. Zoeal stages of Actaea semblatae (Crustacea, Decapoda, Xanthidae), with a key to the known xanthid zoeas of Korea. Korean J. Syst. Zool., 18: 1 12. Lee, D.H., 1993. Larval development of Macromedaeus distinguendus (De Haan, 1835) and Pilumnopeus makiana (Rathbun, 1929) (Brachyura, Xanthidae) reared in the laboratory. Master thesis, Pusan Nat. Univ., pp. 1 53. Lee, K.H. and H.S. Ko, 2008. First records of three crabs (Crustacea: Decapoda) from Korea. Korean J. Syst. Zool., 24: 17 24. Lee, S.H., K.H. Lee and H.S. Ko, 2011. First records of two pilumnid crabs (Crustacea: Decapoda) collected from Jejudo Island, southern Korea. Korean J. Syst. Zool., 27: 191 196. Lee, S.H. and H.S. Ko, 2009a. First record of crinoids symbiotic crab, Harrovia japonica (Decapoda: Brachyura:

66 II Pilumnidae) from Korea. Korean J. Syst. Zool., 25: 125 128. Lee, S.H. and H.S. Ko, 2009b. Zoeal stages of Harrovia japonica (Decapoda: Brachyura: Pilumnidae) with a key to the known eumedoninid zoeae from the Indo-Pacific. Anim. Cells Syst., 13: 213 222. Lee, S.H. and H.S. Ko, 2011 First records of three xanthid crabs (Decapoda: Brachyura: Xanthidae) from Jejudo Island in Korea. Korean J. Syst. Zool., 27: 183 190. Lee, S.K., S.H. Kim and W. Kim, 2008. Report of four species of crabs (Crustacena: Decapoda: Brachyura) from Korea. Korean J. Syst. Zool., 24: 291 297. Lee, S.K., T.S. Park and W. Kim, 2010. New report of two xanthid crabs (Crustacea: Decapoda: Brachyura) from Korea. Korean J. Syst. Zool., 26: 87 91. Linnaeus, C., 1767. Systema Naturae per Regna tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. (edit. 12), 1(2): 533 1327. MacLeay, W.S., 1838. On the Brachyurous Decapod Crustacea brought from the Cape by Dr. Smith. In: Dr. A, Smith, Illustrations of the Annulosa of South Africa; being a Portion of the Objects of Natural History chiefly collected during an Expedition into the interior of South Africa, under the direction of Dr. Andrew Smith, in the years 1834, 1835, and 1836; fitted out by The Cape of Good Hope Association for Exploring Central Africa: 53 71, pls. 2 3. London. Milne Edwards, A., 1865. Études zoologiques sur les Crustacés récents de la famille des Cancériens. Cancérides, Pirimélides, Carpilides, Première Partie. Nouv. Arch. Mus. Nat. Hist. Nat., Paris, 1: 177 308, pls. 11 19. Milne Edwards, A., 1867. Descriptions de quelques espèces nouvelles de Crustacés Brachyures. Ann. Soc. Ent., de France, (ser. 4) 7(4): 263 288. Milne Edwards, A., 1872. Recherches sur la faune carcinologique de la Nouvelle-Calédonie. Part 1. Groupe des Oxyrynches. Nouv. Arch. Mus. Nat. Hist. Nat., Paris, 8: 229 267, pls. 10 14. Milne Edwards, A., 1873. Recherches sur la faune carcinologique de la Nouvelle-Calédonie. Part II. Groupe des Cyclométopes Portuniens. Nouv. Arch. Mus. Nat. Hist. Nat., Paris, 9: 155 332, pls. 4 18. Milne-Edwards, H., 1834. Histoire Naturelle des Crustacés, comprenant l anatomie, la physiologie et la classification de ces animaux. Libraire Encyclopédique de Roret, Paris, Vol. 1: i xxxv, 1 468, pls. 3, 5 6, 15 17, 20, 22 23, 25. Minemizu, R., 2000. Marine Decapod and Stomatopod Crustaceans mainly from Japan. Buninchi Sogo Publishing, Co., Ltd., Tokyo, pp. 1 344 (In Japanese). Miyake, S., 1983. Japanese Crustacean Decapods and Stomatopods in colour. Hoikusha Publ. Co. Ltd., 2: 1 277, pls. 1 64. Ng, P.K.L., 1993. Kraussiinae, a new subfamily for the genera Kraussia Dana, 1852, Palapedia, new genus, and Garthasia, new genus (Crustacea: Decapoda: Brachyura: Xanthidae), with descriptions of two new species from Singapore and the Philippines. Raffles Bull. Zool., 41: 133 157. Ng, P.K.L., D. Guinot and P.J.F. Davie, 2008. Systema Brachyurorum: Part 1. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull. Zool., 17: 1 286. Ng, P.K.L. and M. Takeda, 2010. Zehntneriana, a replacement name for Zehntneria Takeda, 1972 (Crustacea, Brachyura, Pilumnidae), preoccupied by Zehntneria Brunner Von Wattenwyl, 1907 (Insecta, Orthoptera, Phasmidae). Bull. Nat. Sci. Mus., Ser. A (Zool.), Tokyo, 36: 49 50. Odhner, T., 1925. Monographierte Gattungen der Krabben-familie Xanthidae. I. Göteborgs K. Vet. och Vit. Samh. Handl., (4) 29(1): 1 92, figs. 1 7, pls. 1 5. Ortmann, A.E., 1893. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berücksichtigung der von Herrn Dr. Döderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur zeit im Strassburger Museum aufbewahrten Formen. Theil VII. Abteilung Brachyura (Brachyura genuina Boas), II. Unterabteilung: Cancroidea, 2. Section: Cancrinea, 1. Gruppe: Cyclometopa. Zool. Jahr., Abtheilung

67 für Systematik, Geographie und Biologie der Thiere, 7: 411 495, pl. 17. Rathbun, M.J., 1910. The Brachyura. In: The Danish Expedition to Siam, 1899 1900. V. K. Dansk. Vidensk. Selsk. Skr., Kjøbenhavn, 5(4): 301 367, figs. 1 44, pls. 1 2. Rathbun, M.J., 1931. New and rare Chinese crabs. Lingnan Sci. Jour., 8(1929): 75 104, pls. 5 15. Rüppell, E.S., 1830. Beschreibungen und Abbildungen von 24 Arten kurzschwänzigen krabben, als Beiträge zur Naturgeschichte des Rothen Meeres. H.L. Brönner, Frankfurt a. M.: 1 28, pls. 1 6. Sakai, K., 2004. Crabs of Japan. World Biodiversity Database CD-ROM Series. Sakai, T., 1934. Brachyura from the coast of Kyushu, Japan. Science Reports of the Tokyo Bunrika Daigaku, (B) 1(25): 281 330, figs. 1 26, pls. 17 18. Sakai, T., 1935. List of marine animals around Shimoda area. Biol. Rep. Shimoda Mar. Biol. Stat., Tokyo Univ. Liter. Sci., 1: 23 85, pls. 1 3. Sakai, T., 1936. Crabs of Japan: 66 plates in life colours with descriptions. Sanseido, Tokyo (1935): 1 239, figs. 1 122, pls. 1 66. Sakai, T., 1939. Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha, pp. 365 741, figs. 1 129, pls. 42 111, Yokendo Co., Tokyo. Sakai, T., 1965a. On two new genera and five new species of xanthoid crabs from the collection of His Majesty, the Emperor of Japan made in Sagami Bay. Crustaceana, 8: 97 106. Sakai, T., 1965b. The crabs of Sagami Bay, collected by His Majesty the Emperor of Japan, i xvi, 1 206 (English text), figs. 1 27, pls. 1 100: 1 92 (Japanese text): 1 26 (references and index in English): 27 32 (index in Japanese), 1 map. Maruzen Co., Tokyo. Sakai, T., 1976. Crabs of Japan and the adjacent seas. Kodansha Ltd. Japan, pp. 1 773, pls. 1 251. Sakai, T., 1983. Description of a new genera and species of Japanese crabs, together with systematically and biogeographically interesting species. Res. Crust., 12: 1 44, pls. 1 8. Serène, R., 1984. Crustacés Décapodes Brachyoures de I Océan Indien occidental et de la Mer Rouge, Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par Crosnier, A: Carpiliidae et Menippidae. Faune Trop., no. XXIX: 1 243. Stimpson, W., 1858a. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Pars IV. Crustacea Cancroidea et Corystoidea. Proc. Acad. Nat. Sci. Philadelphia, 1858, 10: 31 40. Stimpson, W., 1858b. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars V. Crustacea Ocypodoidea. Proc. Acad. Nat. Sci. Philadelphia, 10: 93 110 (39 56). Stimpson, W., 1907. Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 1856. Smith. Misc. Coll., Washington, 49(1717): 1 240, pls. 1 26. Takeda, M., 1972. Systematic status of Ceratoplax villosa Zehntner and some related species (Crustacea, Decapoda, Brachyura). Proc. Jap. Soc. Syst. Zool., 8: 34 41, fig. 1 3. Takeda, M., 1977. Crabs from the shallow waters off Mage-jima Island, southwest Japan. Bull. Nat. Sci. Mus., Tokyo, Ser. A (Zool.), 3(2): 73 89, figs. 1 15. Takeda, M., 1982. Keys to the Japanese and foreign Crustaceans, Hokuryukan, Toyko, pp. 1 284. Takeda, M., 1990. Zoeal development of five species of xanthid crabs reared in the laboratory. Res. Crust., 18: 23 47. Takeda, M. and H.S. Kim, 1977. A new Actumnus (Crustacea; Decapoda; Brachyura) from Jeju Island, Korea. Korean J. Zool., 30: 135 139.

68 II Takeda, M. and S. Miyake, 1968. Crabs from the East China Sea. I. Corystoidea and Brachygnatha Brachyrhyncha. J. Fac. Agr., Kyushu Univ., 14(4): 541 582, figs. 1 11, pl. 6. Takeda, M. and S. Miyake, 1969a. Pilumnid crabs of the family Xanthidae from the West Pacific. II. Twentyone species of four genera, with descriptions of four new species. OHMU, 2: 93 156, figs. 1 18. Takeda, M. and S. Miyake, 1969b. On two species of the family Xanthidae (Crustacea, Brachyura) from southern Japan. OHMU, 2(9): 195 206, figs. 1 3. Takeda, M. and S. Miyake, 1969c. A new species of the family Goneplacidae (Crustacea, Brachyura) from the Amakusa Islands, Kyushu. Publ. Amakusa Mar. Biol. Lab., 2: 9 15, fig. 1. Tanaka, H. and K. Konishi, 2001. Larval development of the poisonous crab, Atergatis floridus (Linnaeus, 1767) (Crustacea, Decapoda, Xanthidae) described from laboratoey-reared material. Crust. Res., 30: 21 42. Tanaka, H. and K. Konishi, 2009. Atergatis floridus (Linnaeus, 1767) (Decapoda, Xanthidae): re-examination and correction of zoeal characters. Crust. Res., 38: 55 63. Terada, M., 1980. Zoea larvae of four crabs in the subfamily Xanthinae. Zool. Mag., 89: 138 148. Terada, M., 1982. Zoeal development of the menippid crab, Eriphia smithi MacLeay. Zool. Mag., 91: 255 262. Terada, M., 1985. Early zoeal development of the crab, Halimede fragifer De Haan (Xanthidae, Xanthinae). Proc. Jap. Soc. Syst. Zool., 31: 30 37. Terada, M., 1988. Early zoeal development of the crab, Calvactaea tumida Ward (Xanthidae, Zalasiinae). Res. Rep. of Yokosuka High School., 12: 43 50. Ward, M., 1933. New genera and species of marine Decapoda Brachyura from the coasts of New South Wales and Queensland. Austr. Zool., 7(5): 377 394, pls. 21 23. Ward, M., 1936. Crustacea Brachyura from the Coasts of Queensland. Mem. Qld. Mus., 11: 1 13, pls. 1 3. White, A., 1848. Short descriptions of new or little-known Decapod Crustacea. In: November 23, 1847. William Yarrell, Esq., Vice-Presendent, in the Chair. Proc. Zool. Soc. London, 1847(XV): 222 228. Yamaguchi, T., K. Harada, M. Takeda and T. Kikuchi, 1987. Crab fauna of the Amakusa Islands. Calanus, 10: 1 71.

69 1. Eriphia smithii. A. ; B. 1. 2. Sphaerozius nitidus. A. ; B. 1. 3. A. Halimede fragifer, ; B. Echinoecus nipponicus,. 4. Harrovia japonica. A. ; B. 1. 5. Benthopanope indica. A. ; B. 1. 6. Heteropilumnus ciliatus. A. ; B. 1. 7. Pilumnopeus granulatus. A. ; B. 1. 8. Pilumnopeus makianus. A. ; B. 1. 9. Pilumnus minutus. A. ; B. 1. 10. Pilumnus trispinosus. A. ; B. 1. 11. A. Zehntneriana amakusae, ; B. Forestia depressa,. 12. Actaea semblatae. A. ; B. 1. 13. Gaillardiellus orientalis. A. ; B. 1. 14. A. Novactaea pulchella, ; B. Pilodius miersi,. 15. Pilodius nigrocrinitus. A. ; B. 1. 16. A. Madaeops granulosus, ; B. Palapedia integra,. 17. A. Liomera margaritata, ; B. Neoliomera insularis,. 18. Cycloxanthops truncatus. A. ; B. 1. 19. Leptodius exaratus. A. ; B. 1. 20. Macromedaeus distinguendus. A. ; B. 1. 21. Microcassiope orientalis. A, B.. 22. A. Nanocassiope granulipes, ; B. Neoxanthops lineatus,. 23. A. Atergatis floridus, ; B. Atergatis reticulatus,.

70 II 1A 1B

71 2A 2B

72 II 3A 3B

73 4A 4B

74 II 5A 5B

75 6A 6B

76 II 7A 7B

77 8A 8B

78 II 9A 9B

79 10A 10B

80 II 11A 11B

81 12A 12B

82 II 13A 13B

83 14A 14B

84 II 15A 15B

85 16A 16B

86 II 17A 17B

87 18A 18B

88 II 19A 19B

89 20A 20B

90 II 21A 21B

91 22A 22B

92 II 23A 23B