한수지 51(5), , 2018 Note Korean J Fish Aquat Sci 51(5), ,2018 한국부산에서채집된상어류 ( 연골어강 : 판새아강 ) 1 미기록종, Echinorhinus cookei 이우준 김진구 * 부경대학교자원생물학

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1 한수지 51(5), , 2018 Note Korean J Fish Aquat Sci 51(5), ,2018 한국부산에서채집된상어류 ( 연골어강 : 판새아강 ) 1 미기록종, Echinorhinus cookei 이우준 김진구 * 부경대학교자원생물학과 First Reliable Record of Echinorhinus cookei (Chondrichthyes: Elasmobranchii) Collected from Busan, Korea Woo-Jun Lee and Jin-Koo Kim* Department of Marine Biology, Pukyong National University, Busan 48513, Korea One specimen (181.0 cm; TL, total length) of Echinorhinus cookei was collected from Busan, Korea in 2014 using gill net. Echinorhinus cookei was characterized by having no anal fin, no spine at dorsal fin and numerous small dermal denticles at the placoid scale. This species is very similar to the congeneric species, E. brucus, but differed in the shape of placoid scale (star in E. cookei vs. round in E. brucus) and distribution of placoid scale (separated in E. cookei vs. slightly overlapped in E. brucus). We adopted the Korean name, Ga-si-bi-neul-sang-eo, for E. cookei, after Kim and Ryu (2017). Key words: Echinorhinus cookei, First reliable record, Korea 서론 (Ebert et al., 2013; Nelson et al., 2016), (Hatooka et al., 2013), (Choi, 2016; Jeong et al., 2016). Jordan and Metz (1913) 7 [ (Squalus japonicus), (Squalus mitsukurii), (Squatina japonica), (Heterodontus japonicus), (Mustelus manazo), (Triakis scyllium), (Sphyrna zygaena)], Mori (1928), Mori and Uchida (1934), Uchida and Yabe (1939), Mori (1952), Chyung (1977), Choi et al. (1997, 1998), Cho et al. (2014), Kang et al. (2015) (MABIK, 2017). (Kim and Ryu, 2017; MABIK, 2017) Echinorhinus cookei. 재료및방법 Garrick (1960), Compagno (1984), Voight and Weber (2011), vernia caliper 1.0 mm. Compagno (1984), Ebert et al. (2013), Choi (2016). 15% 30 7, 70%. (PKU, Pukyong National University),. 결과및고찰 Family Echinorhinidae (Korean family name: Ga-si-bi-neul-sang-eo-gwa) (Echinorhinidae) (Echinorhinus) 1 (Nelson et al., 2016), m (Compagno, 1984). Korean J Fish Aquat Sci 51(5), , October 2018 This is an Open Access article distributed under the terms of the Creative Commons Attribution Non-Commercial Licens ( which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited. Received 6 August 2018; Revised 20 August 2018; Accepted 23 August 2018 *Corresponding author: Tel: Fax: address: taengko@hanmail.net Copyright 2018 The Korean Society of Fisheries and Aquatic Science 595 pissn: , eissn:

(Echinorhinus) 2, E. brucus, E. cookei. Echinorhinus cookei Pietschmann, 1928 (Korean name: Ga-si-bi-neul-sang-eo) (Fig.

and Yanagisawa, 1983: 465; Compagno, 1984: 27; Chen and Joung, 1993: 35; Last and Stevens, 2009: 43; Fricke et al., 2011: 346; Ebert et al., 2013: 72; Hatooka et al.

2 596 이우준ㆍ김진구,, (Compagno, 1984). Genus Echinorhinus Blainville, 1816 (Korean genus name: Ga-si-bi-neul-sang-eo-sok) Echinorhinus Blainville, 1816: 121 (type species: Squalus spinosus Gmelin, 1789) (Echinorhinus) 2, E. brucus, E. cookei. Echinorhinus cookei Pietschmann, 1928 (Korean name: Ga-si-bi-neul-sang-eo) (Fig. 1) Echinorhinus cookei Pietschmann, 1928: 297 (type locality: Palliser Bay, Cook Strait, New Zealand); Garrick, 1960: 105; Eschmeyer and Herald, 1983: 20; Taniuchi and Yanagisawa, 1983: 465; Compagno, 1984: 27; Chen and Joung, 1993: 35; Last and Stevens, 2009: 43; Fricke et al., 2011: 346; Ebert et al., 2013: 72; Hatooka et al., 2013: 182; Kim and Ryu, 2017: 47; MABIK, 2017: 336. 관찰표본 PKU 10380, 1, 1,810 mm, m, ,,. 기재 %. 11.5%. (Fig. 2A). (spiracle) Fig. 1. Echinorhinus cookei, PKU 10380, female, 1,810 mm in total length, Busan, Korea. Fig. 2. (A) Teeth of Echinorhinus cookei, PKU 10380, (B) Dermal denticle of Echinorhinus cookei, PKU

3 한국산상어류 1 미기록종, Echinorhinus cookei 597. (gill slits) % %, 2 4.6%. 2, , (subterminal notch)., 1 (Fig. 2B). 체색,.. 분포 ( ), (Eschmeyer and Herald, 1983), (Taniuchi and Yanagisawa, 1983), (Chen and Joung, 1993), (Last and Stevens, 2009), (Fricke et al., 2011) 1,100 m (Ebert et al., 2013). Table 1. Comparison of the morphological characters of Echinorhinus cookei Measurements Present study Neotype Garrick (1960) Nontype Taniuchi and Yanagisawa (1983) Number of specimens Total length (mm) In % of total length Trunk at pectoral origin width height Snout tip to outer nostrils mouth eye st gill-opening th gill-opening pectoral origin st dorsal origin nd dorsal origin pelvic origin upper caudal origin Eye horizontal diameter Mouth width length Gill opening lengths 1st th First dorsal fin vertical height length of base Second dorsal fin vertical height length of base Caudal fin length of upper lobe length of lower lobe Pectoral fin length anterior margin Distance from origin to origin of width pectoral and pelvic pelvic and lower caudal origin

4 598 이우준ㆍ김진구 비고 (Echinorhinus cookei) (Echinorhinidae) (Ebert et al., 2013; Nelson et al., 2016),,, (Pietschmann, 1928; Garrick, 1960; Compagno, 1984). Garrick (1960) (neotype) (nontype) Taniuchi and Yanagisawa (1983),,, 5, 1, 2 (Table 1). E. brucus, ( vs. E. brucus ) ( vs. E. burucus ) (Garrick, 1960; Compagno, 1984; Castro, 2011). Echinorhinus cookei Kim and Ryu (2017). 사사 ( ),. ( ). References Blainville HD Prodrome d une nouvelle distribution systématique du règne animal. Bull Soc Philomathique de Paris 8, Castro JI The sharks of North America. Oxford Univ Press, London, U.K. Chen CT and Joung SJ Chondrichthyes. In: Fishes of Taiwan. Shen SC, Lee SC, Shao KT, Mok HK, Chen CT and Chen CH, eds. Depart Zool, Nat Taiwan University, Taipei, Taiwan, Cho HG, Kweon SM and Kim BJ New record of the spadenose Shark, Scoliodon laticaudus (Carcharhiniformes: Carcharhinidae) from south sea, Korea. Korean J Ichthyol 26, Choi Y, Kim IS and Nakaya K New records of Sharks, Sphyrna lewini and Alopias vulpinus (Pisces: Elasmobranchii) in Korea. Anim Syst Evol Divers 13, Choi Y, Kim IS and Nakaya K A taxonomic revision of genus Carcharhinus (Pisces: Elasmobranchii) with description of two new records in Korea. Anim Syst Evol Divers 14, Choi Y Sharks of Korea. MABIK (National Marine Biodiversity Institute of Korea), Seocheon, Koea. Chyung MK The fishes of Korea. Iljisa, Seoul, Korea, Compagno LJV Sharks of the World: an annotated and illustrated catalogue of shark species known to date. FAO Fish Synop 125, Ebert D, Fowler S and Compagno L Sharks of the World. Wild Nature Press, Plymouth, U.K. Eschmeyer WN and Herald ES A field guide to Pacific coast fishes of North America from the Gulf of Alaska to Baja California. Peterson Field Guide Series, 28. Houghton- Mifflin Co., Boston, M.A., U.S.A. Fricke R, Kulbicki M and Wantiez L Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Palaeodivers (Stuttg) 4, Garrick JAF Studies of New Zealand elasmobranchii, part 10. The genus Echinorhinus, with an account of a second species, E. cookei Pietschmann, 1928, from New Zealand waters. Trans R Soc NZ 88, Gmelin JF Amphibia, Pisces. In: Systema Naturae. Caroli a Linné, ed. Leiden, Delamolliere 1, Hatooka K, Yagishita N and Yamaguchi A Echinorhinidae. In: Fishes of Japan with Pictorial Keys to the Species. Nakabo T, ed. Tokai University Press, Kanagawa, Japan. Jeong CH, Choi Y, Oh JK, Kim MS and Jeong SJ An overview of Korean sharks listed by the IUCN and CITES. Korean J Ichthyol 28, Jordan DS and Metz CW A catalogue of the fishes known from the water of Korea. Mem Carnegie Mus 6, Kang CB, Lee WJ, Kim JK and Jung HG A new record of the Pacific sleeper shark, Somniosus pacificus (Squaliformes: Somniosidae) from the western margin of the East Sea, Korea. Korean J Ichthyol 27, Kim JK and Ryu JH Distribution Map of Sea Fishes in Korea. Mapledesign, Busan, Korea. Last PR and Stevens JD Sharks and Rays of Australia, second ed. Harvard University Press, Cambridge, London, U.K. MABIK (National Marine Biodiversity Institute of Korea) National List of Marine Species. Namu Press, Seocheon, Korea, Mori T and Uchida K A revised catalogue of the fishes of Korea. J Soc Chosen Nat Hist 19, 16. Mori T A catalogue of the fishes of Korea. J Pan-Pacific Res Inst 3, 2-8. Mori T Check list of the fishes of Korea. Mem Hyogo Univ Agric 1, Nelson JS, Grande TC and Wilson MVH Fishes of the World. 5th Ed. John Wiley & Sons, Hoboken, Canada, Pietschmann V Neue Fish-arten aus dem Pazifischen Ozean. Anzeiger der Akademie der Wissenschaften, Wien 65,

5 Taniuchi T and Yanagisawa F Occurrence of the prickly shark, Echinorhinus cookei, at Kumanonada, Japan. Jpn J Ichthyol 29, Uchida K and Yabe H The fish-fauna of, Saisyu-to (Quelpart Island) and its adjacent waters. J Soc Chosen Nat Hist 25, Voigt M and Weber D Field guide for sharks of the genus Carcharhinus. Verlag Dr. Friedrich Pfeil, München, Germany, 한국산상어류 1 미기록종, Echinorhinus cookei 599

한수지 50(5), , 2017 Original Article Korean J Fish Aquat Sci 50(5), ,2017 한국산육각복과 (Aracanidae) 어류 1 미기록종, 황줄육각복 (Kentrocapros flavofasciatu

한수지 50(5), , 2017 Original Article Korean J Fish Aquat Sci 50(5), ,2017 한국산육각복과 (Aracanidae) 어류 1 미기록종, 황줄육각복 (Kentrocapros flavofasciatu 한수지 50(5), 589-593, 2017 Original Article Korean J Fish Aquat Sci 50(5),589-593,2017 한국산육각복과 (Aracanidae) 어류 1 미기록종, 황줄육각복 (Kentrocapros flavofasciatus) (Kamohara, 1938) 김경무 김진구 * 부경대학교자원생물학과 New Record