김진일_국문겉표지

대한민국생물지 제 권 호상기문류 절지동물문 : 곤충강 : 딱정벌레목 : 풍뎅이상과

제 권 호 상기문류 절지동물문 : 곤충강 : 딱정벌레목 : 풍뎅이상과 환경부국립생물자원관

제 권 호상기문류 절지동물문 : 곤충강 : 딱정벌레목 : 풍뎅이상과 집필자 : 김진일 성신여자대학교

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,,,,,,. 1992 21. 2010 ABS.., 3 7. 2006. 4 2010 9 158 1,037 16, 7 112 1,163. ( ),,,.. 10.,.

1 3 9 11 25 Family Cetoniidae Leach 27 1. Valgus Scriba 29 2. Nipponovalgus Sawada 29 3. Osmoderma LePeletier and Serville 33 4. Gnorimus LePeletier and Serville 34 5. Trichius Fabricius 36 6. Dicronocephalus Hope 40 7. Rhomborhina Hope 42 8. Pseudotorrynorrhina Mikšić 43 9. Cetonia Fabricius 46 10. Protaetia Burmeister 48 11. Anthracophora Burmeister 55 12. Glycyphana Burmeister 56 13. Gametis Burmeister 58 14. Clinterocera Motschulsky 61 Family Rutelidae MacLeay 62 15. Adoretus Dejean 64 16. Parastasia Westwood 68 17. Popillia Dejean 70 18. Proagopertha Reitter 78 19. Phyllopertha Stephens 80 20. Mimela Kirby 82 21. Callistethus Burmeister 92 22. Blitopertha Reitter 94 23. Anomala Samuelle 99 24. Bifurcanomala Kim 117 25. Chejuanomala Kim 119

2 Family Dynastidae Mac Leay 120 26. Allomyrina Arrow 122 27. Eophileurus Arrow 124 28. Pentodon Hope 125 Family Melolonthidae Leach 127 29. Ectinohoplia Redtenbacher 129 30. Hoplia Illiger 131 31. Apogonia Kirby 137 32. Brahmina Blanchardt 140 33. Lasiopsis Erichson 144 34. Pseudosymmchia Dalla Torre 146 35. Miridiba Reitter 148 36. Sophrops Fairmaire 149 37. Holotrichia Hope 153 38. Heptophylla Motschulsky 166 39. Hilyotrogus Fairmaire 167 40. Polyphylla Harris 169 41. Melolontha Fabricius 171 42. Gastroserica Brenske 176 43. Paraserica Reitter 178 44. Maladera Mulsant 179 45. Sericania Motschulsky 200 46. Nipponoserica Nomura 206 47. Serica Mac Leay 209 214 227 258 261

3 Class Insecta Order Coleoptera Scarabaeoidea Latreille, 1802 Pleurosticti Family Cetoniidae Leach, 1815 Subfamily Valginae Mulsant, 1842 Genus Valgus Scriba, 1790 Valgus koreanus Sawada, 1944 Genus Nipponovalgus Sawada, 1941 Nipponovalgus angusticollis (Waterhouse), 1875 Subfamily Trichiinae Fleming, 1821 Genus Osmoderma Le Peletier de Saint-Fargeau and Serville, 1828 Osmoderma opicum Lewis, 1887 Genus Gnorimus Le Peletier de Saint-Fargeau and Serville, 1828 Gnorimus subopacus Motschulsky, 1860 Genus Lasiotrichius Reitter, 1899 Lasiotrichius succinctus (Pallas), 1781 Subfamily Cetoniinae Leach, 1815 Tribe Phaedimini Schoch, 1894 Genus Dicranocephalus Hope, 1831 Dicronocephalus adamsi Pascoe, 1863 Tribe Coryphocerina Burmeister, 1842 Genus Rhomborrhina Hope, 1837 Rhomborrhina polita Waterhouse, 1875 Genus Pseudotorynorrhina Mikšić, 1974 Pseudotorynorrhina japonica (Hope), 1841 Tribe Cetoniini Leach, 1815 Genus Cetonia Fabricius, 1775 Cetonia pilifera (Motschulsky), 1860 Cetonia pilifera magnifica (Ballion) Genus Protaetia Burmeister, 1842 Protaetia lugubris (Herbst), 1786 Protaetia brevitarsis seulensis (Kolbe), 1886 Protaetia famelica scheini (Mikšić), 1959

4 Protaetia mandschuriensis (Schurhoff), 1933 Protaetia orientalis submarmorea (Burmeister), 1842 Genus Anthracophora Burmeister, 1842 Anthracophora rusticola Burmister, 1842 Genus Glycyphana Burmeister, 1842 Glycyphana fulvistemma Motschulsky, 1860 Genus Gametis Burmeister, 1842 Gametis jucunda (Faldermann), 1935 Tribe Cremastocheilini Burmeister and Schaum, 1841 Genus Clinterocera Motschulsky, 1858 Clinterocera obsoleta (Fairmaire), 1878 Family Rutelidae MacLeay, 1819 Tribe Adoretini Burmeister, 1844 Genus Adoretus Dejean, 1833 Adoretus tenuimaculatus Waterhouse, 1875 Adoretus hirsutus Ohaus, 1914 Tribe Rutelini MacLeay, 1819 Genus Parastasia Westwood, 1842 Parastasia ferrieri Nonfried, 1895 Tribe Anomalini Streubel, 1839 Subtribe Popillina Ohaus, 1918 Genus Popillia Dejean, 1821 Popillia flavosellata Fairmaire, 1886 Popillia mutans Newman, 1838 Popillia quadriguttata (Fabricius), 1787 Popillia quadriguttata japonica Newmann Popillia formosana Arrow, 1913 Subtribe Anomalina Streubel, 1839 Genus Proagopertha Reitter, 1903 Proagopertha lucidula (Faldermann), 1935 Genus Phyllopertha Stephens, 1830 Phyllopertha diversa Waterhouse, 1875 Phyllopertha horticola (Linne), 1758 Genus Mimela Kirby, 1823 Mimela fusania Bates, 1888 Mimela splendens (Gyllenhal), 1817 Mimela holosericea (Fabricius), 1787 Mimela testaceipes (Motschulsky), 1860

5 Mimela pekinensis coreana Machatschke, 1952 Mimela costata (Hope), 1839 Genus Callistethus Blanchard, 1851 Callistethus plagiicollis (Fairmaire), 1886 Genus Blitopertha Reitter, 1903 Blitopertha conspurcata (Harold), 1878 Blitopertha pallidipennis Reitter, 1903 Blitopertha orientalis (Waterhouse), 1875 Genus Anomala Samuelle, 1819 Anomala sieversi Heyden, 1887 Anomala albopilosa Hope, 1839 Anomala japonica Arrow, 1913 Anomala mongolica (Faldermann), 1835 Anomala geniculata (Motschulsky), 1866 Anomala chamaeleon Fairmaire, 1887 Anomala rufocuprea Motschulsky, 1860 Anomala koreana Kim, 1997 Anomala luculenta Erichson, 1847 Anomala corpulenta Motschulsky, 1853 Anomala ignicolor Reitter, 1903 Anomala ogloblini S. Medvedev, 1949 Anomala cuprea Hope, 1839 Genus Bifurcanomala Kim, 1998b Bifurcanomala aulax (Wiedemann), 1923 Genus Chejuanomala Kim, 1998b Chejuanomala quelparta (Okamoto), 1924 Family Dynastidae MacLeay, 1819 Tribe Dynastini MacLeay, 1819 Genus Allomyrina Arrow, 1911 Allomyrina dichotoma (Linne), 1771 Tribe Phileurini Burmeister, 1847 Genus Eophileurus Arrow, 1908 Eophileurus chinensis (Faldermann), 1835 Tribe Pentodontini Mulsant, 1842 Genus Pentodon Hope, 1837 Pentodon quadridens (Gebler), 1845 Family Melolonthidae Leach in Samouelle, 1819 Subfamily Hoplinae Latreille, 1829

6 Genus Ectinohoplia Redtenbacher, 1868 Ectinohoplia rufipes (Motschulsky), 1860 Genus Hoplia Illiger, 1803 Hoplia aureola (Pallas), 1871 Hoplia djukini Jacobson, 1914 Hoplia koreana Moser Subfamily Melolonthinae Leach in Samouelle, 1819 Tribe Rhizotrogini Burmeister, 1855 Genus Apogonia Kirby, 1818 Apogonia cribricollis Burmeister, 1855 Apogonia cupreoviridis Kolbe, 1886 Genus Brahmina Blanchardt, 1850 Brahmina rubetra faldermanni (Murayama), 1954 Brahmina intermedia (Mannerheim), 1849 Brahmina darcis Reitter, 1902 Brahmina crenicollis (Motschulsky) Genus Lasiopsis Erichson, 1848 Lasiopsis sahlbergi (Mannerheim), 1949 Lasiopsis manchuricus Murayama, 1941 Genus Pseudosymmchia Dalla Torre, 1913 Pseudosymmchia impressifrons (Fairmaire), 1887 Genus Miridiba Reitter, 1902 Miridiba castanea (Waterhouse), 1875 Genus Sophrops Fairmaire, 1887 Sophrops heydeni (Brenske), 1982 Sophrops striata (Brenske), 1892 Genus Holotrichia Hope, 1837 Holotrichia niponensis (Lewis), 1985 Holotrichia parallela (Motschulsky), 1854 Holotrichia koraiensis Murayama, 1937a Holotrichia picea (Waterhouse), 1875 Holotrichia ernesti Reitter, 1902 Holotrichia diomphalia (Bates), 1888 Holotrichia oblita (Faldermann), 1835 Holotrichia reticulata Murayama, 1941 Genus Heptophylla Motschulsky, 1858 Heptophylla picea Motschulsky, 1857 Genus Hilyotrogus Fairmaire 1886

7 Hilyotrogus bicoloreus (Heyden), 1887 Holotrichia kiotoensis Brenske Holotrichia sichotana Brenske Tribe Melolonthini Leach in Samouelle, 1819 Genus Polyphylla Harris, 1841 Polyphylla laticollis manchurica Semenov, 1900 Genus Melolontha Fabricius, 1775 Melolontha incana (Motschulsky), 1853 Melolontha insulana (Moser), 1918 Subfamily Sericinae Kirby, 1837 Genus Gastroserica Brenske, 1897b Gastroserica herzi (Heyden), 1887 Genus Paraserica Reitter, 1896 Paraserica grisea (Motschulsky), 1866 Genus Maladera Mulsant and Rey, 1871 Maladera holosericea (Scopoli), 1772 Maladera coreana Kim and Kim, 2003 Maladera verticalis (Fairmaire), 1888 Maladera castanea koreana Kim and Kim, 2003 Maladera ovatula (Fairmaire), 1891 Maladera aureola (Murayama), 1938 Maladera renardi (Ballion), 1870 Maladera schoenfeldti (Murayama), 1937 Maladera gibbiventris (Brenske), 1897 Maladera infuscata (Moser), 1915 Maladera fusania (Murayama), 1934 Maladera opaciventris (Moser), 1915 Maladera orientalis (Motschulsky), 1857 Maladera cariniceps (Moser), 1915 Genus Sericania Motschulsky, 1860 Sericania fuscolineata Motschulsky, 1860 Sericania latisulcata Murayama, 1941 Sericania koryoensis Murayama, 1935 Sericania yamauchii Sawada, 1938 Genus Nipponoserica Nomura, 1973 Nipponoserica elliptica (Murayama), 1938 Nipponoserica opacicarina Kim and Kim, 2003 Genus Serica MacLeay, 1819

8 Serica fulvopubens (Reitter), 1986 Serica lutea Kim and Kim, 2003 Serica septentrionalis Murayama, 1935 Serica hirsuta Kim and Kim, 2003

9 (Coleoptera) (Scarabaeoidea) 3 7 ( ). ( ),, (, ) 5-5-5 ( ). ( ). Lamellicornia (Lucanidae) (Passalidae). (Hydrophilidae) (Gyrinidae),. 2,500 35,000,,. Lawrence and Newton (1995) 1, 13 90% (Scarabaeidae, sensu lato).. (, pleural membrane) (, Laparosticti) (, Pleurosticti), (Scarabaeidae), (Aphodiidae) 10, ( ) (Melolonthidae), (Dynastidae), (Rutelidae), (Cetoniidae)., (Trogidae), (Geotrupidae).,,,.,,.,,,,. (Polyphaga) (urogomphi: ), Crowson (1955, 1960) (Staphyliniformia) Dascillidae (Elateriformia). Vogler Michael (Michael et al.,, 2005) Hansen (1995, 1997). : Crowson (1955, 1960), Lawrence and Newton (1995), Hansen (1995, 1997), Michael et al. (2005), Andrew and Smith (2006). : Kolbe (1886) 3 10, 4 18 28 Heyden (1887) 9 5. 13 (Lawrence 7 ) 400,,

10 9 250. (Kim, 2000, 2001b) 13 234,. 4 109 4 113. (Kim, 2000, 2001b). : Kolbe (1886), Heyden (1887), Bates (1888), Kraatz (1888), Reitter (1895), Murayama (1935b, 1937a, 1954), Cho (1969), Stebnicka (1980), Kim (1992, 1993, 2000, 2001b).

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13 Table 1. Species names of Scarabaeoidea Pleurosticti recorded from Korea (2009). Reported generic names Specific names 1 1 Valgus Dasyvalgus Nipponovalgus angusticollis Valgus koreanus 2 Valgus Chromovalgus Charitovalgus fumosus 3 Valgus pictus fumosus 4 5 Valgus hemipterus 5 Osmoderma barnabita 6 6 Osmoderma caeleste 7 Osmoderma apica 8 7 Osmoderma opicum 9 Gnorimus subopacus 10 8 Gnorimus subopacus/subcostatus 11 Trichius Lasiotrichius succinctus 12 10 Trichius abdominalis 13 11 Paratrichius Gnorimus septemdecimguttatus 14 12 Dicronocephalus adamsi 15 13 Rhomborhina Torynorhina polita 16 14 Rhomborhina Torynorhina japonica/coreana 17 Rhomborhina unicolor 18 Rhomborhina nigra 19 15 Dasyvalgus Rhomborhina japonica 20 16 Glycyphana Cetonia (Eucetonia) 17 Eucetonia Chromovalgus pilifera 21 pilifera pilifera 22 pilifera magnifica 23 aurata var. viridiopaca 24 aurata var. cupreola 25 aurata var. pilifera 26 magnifica 27 roelofsi 28 viridiopaca 29 Potosia (Potosia) metallica daurica 30 Cetonia seulensis 31 18 Glycyphana cupreola 32 19 Protaetia lugubris 33 20 Potosia (Liocola) Nectosia (Liocola) lugubris orientalis 34 Cetonia seulensis 35 16 Cetonia 22 Liocola Potosia (Liocola) 18 Protaetia brevitarsis 36 23 Nectosia (Liocola) 23 Pachnotosia Protaetia Liocola brevitarsis seulensis 37 Liocola brevitarsis viridis 38 22 Liocola Protaetia Cetonia speculifera 39 19 Protaetia cataphracta 40 Protaetia pryeri 41

14 Table 1. Continued. Reported generic names Specific names Potosia famelica/amurensis 42 Potosia amurensis 43 Potosia (Potosia) Protaetia Nectosia (Potosia) famelica 44 Protaetia famelica scheini 45 Protaetia mandschuriensis 46 Cetonia Potosia (Calopotosia) submarmorea 47 Protaetia orientalis submarmorea 48 Potosia Potosia (Calopotosia) aerata 49 Protaetia orientalis 50 Protaetia ishigakia 51 Protaetia lewisi 52 24 Anthracophora 25 Poecillophilides rusticola 53 26 Glycyphana fulvistemma 54 Glycyphana kuperi 55 Glycyphana albosetosa 56 Glycyphana argyroticta 57 Glycyphana 27 Oxycetonia 28 Gametis jucunda 58 27 Oxycetonia jucunda ferruginosa Oxycetonia jucunda borealis Oxycetonia jucunda dolens Oxycetonia jucunda bealiae Oxycetonia jucunda kuperi Oxycetonia jucunda albosetosa Oxycetonia jucunda argyrosticta 29 Clinterocera 30 Callynomes obsoleta 59 Clinterocera mandarina 60 Cetoniidae 30 genera 60 sp. names+10 ssp. names 1 Parastasia ferrieri 1 2 Ohkubous ferrieri quadridentatus 2 3 Adoretus falciungulatus 3 Adoretus hirsutus 4 Adoretus sinicus 5 Adoretus tenuimaculatus 6 Adoretus umbrosus tenuimaculatus 7 4 Popillia/Popilia atroco(a)erulea 8 Popillia/Popilia atrocoerulea atra Popillia/Popilia atrocoerulea biplagiata Popillia/Popilia atrocoerulea loczyi Popillia/Popilia coerulea 9 Popillia adamas 10

15 Table 1. Continued. Reported generic names Specific names Popillia flavosellata 11 5 Popilia/Popillia bogdanowi 12 Popillia chinensis/sordida 13 Popillia formosana 14 Popilia/Popillia indigonacea 15 Popillia/Popilia mutans 16 Popillia/Popilia mutans indigonacea Popilia/Popillia japonica/bogdanowi 17 Popillia quadriguttata 18 Popillia quadrig. quadriguttata Popillia quadriguttata japonica Popillia pallipennis 19 Popillia quelpartiana 20 Popillia ruficollis 21 Popillia uchidai 22 6 Proagopertha 7 Phyllopertha 8 Rhombonyx lucidula 23 Proagopertha Phyllopertha Ph.(Proagopertha) pubicollis 24 9 Phyllopertha diversa 25 Phyllopertha diversa koreana Phyllopertha horticola 26 Phyllopertha intermixta 27 10 Blitopertha Phyllopertha conspu(e)rcata 28 Blitopertha 11 Anomala Blitopertha (Exomala) Phyllopertha orientalis 29 Blitopertha Phyllopertha Blitopertha (Exomala) pallidipennis/nigrata 30 12 Mimela chinensis 31 Mimela concolor 32 Mimela coreana Niijima and K. 33 Mimela Anomala costata 34 Mimela Anomala difficilis 35 Mimela flavilabris 36 Mimela fusania 37 Mimela Anomala Rhombonyx holosericeus 38 Mimela lucidula 39 Mimela lucidula gaschkewitschi Mimela lucidula coruscus Mimela Mim.(Paramimela) pekinensis c(k)oreana 40 Mimela splendens 41 Mimela splendens cyanicollis Mimela splendens corusca Mimela testaceoviridis 42 Mimela Anomala Mim.(Paramimela) Rhombonyx testaceipes 43

16 Table 1. Continued. Reported generic names Specific names 11 Anomala testaceipes ochroptera 8 Rhombonyx testaceipes ussuriensis 12 Spilota Anomala Anomala (Spilota) A. (Anomala) plagiicollis 44 13 Callistethus plagiicollis 14 Anomala (Paraspilota) A. (Anomala) Anomala (Spilota) impicta 45 Anomala (Spilota) plagiicollis impicta 46 Anomala A. (Euchoronomala) albopilosa 47 Anomala Anomala (Emphalena) chamaeleon 48 Anomala corpulenta 49 Anomala corrugata 50 Anomala A. (Anomala) A. (Idiocnema) costifera 51 Anomala 14 Euchlora A. (Euchoronomala) A. (Aprosterna) cuprea 52 Anomala daimiana 53 Anomala daurica rufofemorata 54 Anomala dubia 55 Anomala geniculata 56 Anomala gottschei 57 Anomala ignicolor 58 Anomala japonica 59 Anomala koreana 60 Anomala luculenta 61 Anomala luculenta cuprea Anomala luculenta purpurea Anomala luculenta smaragdina Anomala lucens 62 Anomala A. (Euchlora) A. (Euchoronomala) mongolica 63 Anomala A. (Aprosterna) motschulskyi 64 Anomala motsch. viridicuprea Anomala Heteroplia multistriata 65 Anomala octiescostata 66 Anomala A. (Chrysoplethisa) A. (Phyllopertha) octocostata 67 Anomala (Anomala) ogloblini 68 Anomala orientalis 69 Anomala Anomala (Anomala) pleurimago 70 Anomala Anomala (Anomala) polyanor 71 Anomala puncticollis 72 Anomala rufocuprea 73 Anomala schoenfeldti 74 Anomala Phyllopertha A. (Euporomala) sieversi 75 Anomala sieversi atrocoerulea Anomala sieversi octocostata

17 Table 1. Continued. Reported generic names Specific names Anomala sieversi subpurpurea Anomala A. (Idiocnema) spiloptera 76 Anomala A. (Euchlora) A.(Euchoronomala) viridana 77 15 Euchlora cuprea viridana 78 16 Bifurcanomala Anomala aulax 79 17 Chejuanomala Anomala quelparta 80 Rutelidae 17 genera 80 names+20 ssp. Names 3 Allomyrina Trypoxylus Xylotrupes dichotoma 1 5 Eophileurus Phileurus chinensis 2 Phileurus morio 3 6 Pentodon quadridens 4 7 Oryctes rhinoceros 5 Dynastidae 7 genera 5 names 1 Ectinohoplia Hoplia rufipes 1 Ectinohoplia obducta 2 2 Hoplia 12-punctata 3 Hoplia H. (Euchromoplia) aureola/aureula 4 Hoplia Hoplia (Decamera) djukini 5 Hoplia communis 6 Hoplia koreana 7 Hoplia moerens 8 Hoplia squamacea 9 3 Apogonia cribricollis 10 Apogonia amida 11 Apogonia chinensis 12 Apogonia cupreoviridis 13 Apogonia cupreoviridis fusania 14 Apogonia nigroolivacea 15 4 Brahmina rubetra/faldermanni 16 Brahmina faldermanni 17 Brahmina intermedia/sibirica 18 Brahmina darcis 19 Brahmina crenicollis 20 Brahmina excisiceps 21 5 Cyphochilus farinosus 22 6 Lasiopsis sahlbergi 23 Lasiopsis formosanus 24 Lasiopsis manchuricus 25 7 Pseudosymmchia 8 Metabolus impressifrons 26 9 Rhizotrogus 10 Holotrichia 11 L. (Holotrichia) Lachnosterna niponicus 27

18 Table 1. Continued. Reported generic names Specific names 12 Miridiva 13 Miridiba koreana 28 Miridiva coreana 29 Miridiba Holotrichia Holotrich. (Pledina) Miridiva castanea 30 14 Neodontocnema Lachnosterna L.(Holotrichia) Sophrops heydeni 31 Brahmina heydeni 15 Sophrops striata 32 11 Lachnosterna Holotrichia niponensis 33 10 Holotrichia 17 Eotrichia Lachnosterna titanis 34 Lachnosterna Holotrichia parallela 35 Lachnosterna Holotrichia morosa 36 Lachnosterna Holotrichia inelegans 37 Holotrichia koraiensis 38 Lachnosterna Holotrichia picea 39 Holotrichia ernesti 40 Lachnosterna Holotrichia diomphal(r)ia 41 Holotrichia oblita 42 16 Holotrichia Pollaplonyx (Bunbunius) reticulata 43 Holotrichia Lachnosterna kiotoensis 44 Holotrichia sichotana 45 Holotrichia convexopyga 46 Holotrichia novila 47 Lachnosterna Holotrichia picea 48 17 Heptophylla picea 49 18 L.(Holotrichia) Hilyotrogus bicoloreus 50 19 Polyphylla laticollis 51 Polyphylla laticollis manchurica 52 Polyphylla laticollis mandshurica Polyphylla laticollis mandschurica Polyphylla laticollis mandsurica Polyphylla chinensis manchurica 53 20 Melolontha 20 Hoplosternus incana/incanus 54 Melolontha insulana 55 Melolontha frater 56 Melolontha japonica 57 21 Hoplosternus japonicus 58 16 Pollaplonyx flavidus 59 22 Serica Gastroserica 24 Microserica herzi/hertzi 60 25 Paraserica Serica grisea 61 26 Aserica Serica 27 Maladera aureola 62 28 Trichoserica Serica 29 Ophthalmoserica boops 63 Serica brunnea 64

19 Table 1. Continued. Reported generic names Specific names 30 Autoserica Aserica Serica Maladera cariniceps 65 Autoserica Serica Aserica Maladera castanea 66 Maladera (Aserica) castanea Aserica Serica Maladera (Maladera) fusania 67 Autoserica Serca Aserca Maladera gibbiventris 68 27 Maladera Serica holosericea (h.-rica) 69 Maladera coreana 70 Maladera Mal. (Eumaladera) nitidiceps 71 Maladera Serica Aserica verticalis 72 Maladera castanea koreana 73 Autoserica Serica Aserica Maladera infuscata 74 Maladera (Aserica) infuscata Autoserica Aserica Serica Maladera ovatula/ovatura 75 Autoserica Serica Autos. (Maladera) Maladera japonica 76 Autoserica Aserica Serica Maladera koreana 77 Maladera Serica renardi 78 Serica Maladera (Aserica) motschulskyi 79 Serica salebrosa 80 Serica Maladera Maladera (Aserica) spissigrada 81 Serica nakayamai 82 Serica formosana 83 Maladera Serica schoenfeldti 84 Maladera Serica Aserica Maladera secreta 85 Aserica Serica Maladera okamotoi 86 Aserica Serica Maladera fusania 87 Autoserica Serica Aserica Maladera opaciventris 88 Serica Aserica Maladera orientalis 89 Maladera kamiyai 90 Maladera Serica thibetana 91 Maladera Serica Aserica laboriosa 92 Maladera Serica Aserica stridula 93 Maladera Aserica Autoserica Serica formosae 94 30 Sericania Serica fuscolineata 95 Sericania Serica latisulcata 96 Sericania Serica koryoensis 97 Sericania Serica yamauchii 98 Sericania hasegawai 99 Sericania Maladera kamiyai 100 Sericania angulata 101 Sericania imadatei 102 31 Nipponoserica Serica elliptica 103

20 Table 1. Continued. Reported generic names Specific names Nipponoserica opacicarina 104 Nipponoserica Serica similis 105 32 Trichoserica Serica 33 Omaloplia polita 106 Trichoserica fulvopubens 107 Serica lutea 108 Serica Trichoserica septentrionalis 109 Serica hirsuta 110 Serica nakayamai 111 Serica Maladera (Aserica) motschulskyi 112 Ophthalmoserica sawadai 113 Serica Ophthalmoserica formosana 114 Melolonthidae 33 genera 114 names Total: 4 families 87 genera 289 specific and subspecific names

21 Table 2. Species list of Korean pleurosticti scarabs, with the observed number of specimens, and with doubtful and removed species from the Korean fauna (2009). T st Registrated Species Indiv. Doubtful+Removed sp. 1 1 Valgus koreanus Sawada 25 Valgus hemipterus 2 2 Nipponovalgus angusticollis (Waterhouse) 187 Charitovalgus fumosus 3 3 Osmoderma opicum Lewis 5 Osmoderma barnabita 4 4 Gnorimus subopacus Motschulsky 179 Osmoderma caeleste 5 5 Trichius succinctus (Pallas) 2,597 Osmoderma apica/opicum 6 6 Dicronocephalus adamsi Pascoe 307 Gnorimus subcostatus 7 7 Rhomborhina polita Waterhouse 3 Paratrichius septemdecimguttatus 8 8 Pseudotorynorhina japonica (Hope) 310 Trichius abdominalis 9 9 Cetonia pilifera p./p. magnifica 562 Rhomborhina nigra 10 10 Protaetia lugubris (Herbst) 21 Rhombonyx ussuriensis 11 11 Protaetia brevitarsis seulensis (Kolbe) 472 Torynorhyna unicolor 12 12 Protaetia famelica scheini (Mikšić) 97 Cetonia roelofsi 13 13 Protaetia mandschuriensis (Schürhoff) 91 Cetonia viridiopaca 14 14 P. orientalis submarmorea (Burmeister) 745 Cetonia aurata viridiopaca 15 15 Anthracophora rusticola Burmister 539 Cetonia/(Potosia) metallica dauria 16 16 Glycyphana fulvistemma Motschulsky 342 Potosia aerata 17 17 Gametis jucunda (Faldermann) 5,358 Potosia amurensis 18 18 Clinterocera obsoleta Fairmaire 201 Potosia lugubris orientalis Protaetia cataphracta Protaetia cuprea daurica Protaetia ishigakia Protaetia lewisi Protaetia pryeri Protaetia speculifera Liocola speculifera Glycyphana cupreola Glycyphana kuperi Clinterocera mandarina Cetoniidae: 14 gen. 18 spp. 11,627ex. 15 gen. 28 spp. 19 1 Adoretus tenuimaculatus Waterhouse 3,280 Adoretus falciungulatus 20 2 Adoretus hirsutus Ohaus 214 Adoretus sinicus 21 3 Parastasia ferrieri Nonfried 13 Popillia atrocoerulea 22 4 Popillia flavosellata Fairmaire 2,450 Popillia biguttata 23 5 Popillia mutans Newman 2,079 Popillia chinensis 24 6 Popillia quadriguttata (Fabricius) 875 Popillia indigonacea 25 7 Popillia formosana Arrow 19 Popillia quelpartiana 26 8 Proagopertha lucidula (Faldermann) 236 Popillia uchidai 27 9 Phyllopertha diversa Waterhouse 557 Popilia adamas 28 10 Phyllopertha horticola (Linné) 3 Proagopertha pubicollis

22 Table 2. Continued. T st Registrated Species Indiv. Doubtful+Removed sp. 29 11 Mimela fusania Bates 732 Phyllopertha intermixta 30 12 Mimela splendens (Gyllenhal) 1,787 Phyllopertha octiescostata 31 13 Mimela holosericea (Fabricius) 124 Mimela chinensis 32 14 Mimela testaceipes (Motschulsky) 1,117 Mimela concolor 33 15 M. pekinensis coreana Machatschke 0 Mimela coreana 34 16 Mimela costata (Hope) 1 Mimela costata 35 17 Callistethus plagiicollis (Fairmaire) 1,057 Mimela difficilis 36 18 Blitopertha conspurcata (Harold) 1,464 Mimela flavilabris 37 19 Blitopertha pallidipennis Reitter 2,686 Mimela testaceoviridis 38 20 Blitopertha orientalis (Waterhouse) 439 Anomala corrugata 39 21 Anomala sieversi Heyden 1,175 Anomala costifera 40 22 Anomala albopilosa Hope 835 Anomala cuprea 41 23 Anomala japonica Arrow 130 Anomala daimiana 42 24 Anomala mongolica (Faldermann) 1,057 Anomala daurica rufofemorata 43 25 Anomala geniculata (Motschulsky) 1 Anomala lucens 44 26 Anomala chamaeleon Fairmaire 3,941 Anomala motschulskyi 45 27 Anomala rufocuprea Motschulsky 362 Anomala multistriata 46 28 Anomala koreana Kim 25 Anomala orientalis 47 29 Anomala luculenta Erichson 684 Anomala pleurimago 48 30 Anomala corpulenta Motschulsky 1,033 Anomala polyanor 49 31 Anomala ignicolor Reitter 1 Anomala puncticollis 50 32 Anomala ogloblini S. Medvedev 0 Anomala schoenfeldti 51 33 Anomala cuprea Hope 0 Anomala spiloptera 52 34 Bifurcanomala aulax (Wiedemann) 781 Anomala viridana 53 35 Chejuanomala quelparta (Okamoto) 138 Rutelidae: 11 gen. 35 spp. 29,297ex 6 gen. 34 spp. 54 1 Allomyrina dichotoma (Linné) 51 Oryctes rhinoceros 55 2 Eophileurus chinensis (Faldermann) 72 Phileurus morio 56 3 Pentodon quadridens (Gebler) 6 Dynastidae: 3 gen. 3 spp. 129ex 2 gen. 2 spp. 57 1 Ectinohoplia rufipes (Motschulsky) 774 Ectinohoplia obducta 58 2 Hoplia aureola (Pallas) 243 Hoplia communis 59 3 Hoplia (D.) djukini Jacobson 0 Hoplia moerens x x Hoplia koreana: no given species number Hoplia squamacea 60 4 Apogonia cribricollis Burmeister 64 Apogonia amida 61 5 Apogonia cupreoviridis Kolbe 544 Apogonia chinensis 62 6 Brahmina rubetra faldermanni (Muray.) 51 Apogonia nigroolivacea 63 7 Brahmina intermedia (Mannerheim) 0 Brahmina crenicollis 64 8 Brahmina darcis Reitter 0 Brahmina excissiceps x x Brahmina crenicollis: no given species number

23 Table 2. Continued. T st Registrated Species Indiv. Doubtful+Removed sp. 65 9 Lasiopsis sahlbergi (Mannerheim) 12 66 10 Lasiopsis manchuricus Murayama 0 67 11 Pseudosymmchia impressifrons (Fairmaire) 847 Cyphochilus farinosus 68 12 Miridiba castanea (Waterhouse) 3 69 13 Sophrops heydeni (Brenske) 153 70 14 Sophrops striata (Brenske) 962 71 15 Holotrichia niponensis (Lewis) 216 Holotrichia convexopyga 72 16 Holotrichia parallela (Motschulsky) 1,228 Holotrichia kiotoensis 73 17 Holotrichia koraiensis Murayama 57 Holotrichia morosa 74 18 Holotrichia picea (Waterhouse) 23 Holotrichia novila 75 19 Holotrichia ernesti Reitter 24 Holotrichia sichotana 76 20 Holotrichia diomphalia (Bates) 2,456 77 21 Holotrichia oblita (Faldermann) 2 78 22 Holotrichia reticulata Murayama 20 x x Holotrichia kiotoensis: no given species number x x Holotrichia sichotana: no given species number 79 23 Heptophylla picea Motschulsky 12 80 24 Hilyotrogus bicoloreus (Heyden) 507 81 25 Polyphylla laticollis manchurica Semenov 73 Polyphylla chinensis 82 26 Melolontha incana (Motschulsky) 458 Melolontha frater 83 27 Melolontha insulana (Moser) 1 Melolontha japonica 84 28 Gastroserica herzi (Heyden) 1,604 85 29 Paraserica grisea (Motschulsky) 0 Pollaplonyx flavidus 86 30 Maladera holosericea (Scopoli) 149 Maladera formosae 87 31 Maladera coreana Kim and Kim 19 Maladera japonica 88 32 Maladera verticalis (Fairmaire) 943 Maladera laboriosa 89 33 M. castanea koreana Kim and Kim 53 Maladera motschulskyi 90 34 Maladera ovatula (Fairmaire) 161 Maladera nitidiceps 91 35 Maladera aureola (Murayama) 17 Maladera okamotoi 92 36 Maladera renardi (Ballion) 71 Maladera secreta 93 37 Maladera schoenfeldti (Murayama) 88 Maladera similis 94 38 Maladera gibbiventris (Brenske) 146 Maladera stridula 95 39 Maladera infuscata (Moser) 121 Maladera thibetana 96 40 Maladera fusania (Murayama) 805 Ophthalmoserica sawadai 97 41 Maladera opaciventris (Moser) 0 98 42 Maladera orientalis (Motschulsky) 954 99 43 Maladera cariniceps (Moser) 577 100 44 Sericania fuscolineata Motschulsky 525 Sericania angulata 101 45 Sericania latisulcata Murayama 8 Sericania hasegawai 102 46 Sericania koryoensis Murayama 105 Sericania imadatei 103 47 Sericania yamauchii Sawada 35 Sericania kamiyai

24 Table 2. Continued. T st Registrated Species Indiv. Doubtful+Removed sp. 104 48 Nipponoserica elliptica (Murayama) 40 Nipponoserica similis 105 49 Nipponoserica opacicarina Kim and Kim 22 Serica boops 106 50 Serica polita (Gebler) 172 Serica brunnea 107 51 Serica lutea Kim and Kim 2 Serica formosana 108 52 Serica septentrionalis Murayama 0 Serica koreana 109 53 Serica hirsuta Kim and Kim 53 Serica sawadai Melolonthidae: 17 gen. 53 spp. 15,404ex 14 gen. 41 spp Total 4 families 45 genera 119+4 species 56,457 individuals and with doubtful sp. and removed sp. names 104 (37 genera) *T: species total of 4 subfamilies, st: subtotal of each subfamily. X: no counted Korean species.

25 Class Insecta Order Coleoptera Scarabaeoidea Latreille, 1802 ( ) 5 6. ( ). 3 7 ( )., 5.,. 25 35,000. 250 13,. 1.. 2. 3 2., (geniculate form)., Lucanidae., Passalidae 3. 3,., 2 Laparosticti 4 3,., 2 Pleurosticti 10 4.. ( ) 5.

26, (setal group). Trogidae. 6., 5 5. 8 9.., (hind tibial spur) 1. Scarabaeidae 9 11,., 2 6 6.. 9.. Aphodiidae 7 7. 9.,. Aegialiidae 10. 8 8. 11,. Geotrupidae 10 9 9.. Hybosoridae.. Ochodaeidae 10.,. 2 Dynastidae 11 11. 2, 2., Rutelidae 2, 12 12.. Melolonthidae., Cetoniidae Pleurosticti 2009 4 1. 87, 289.

: : 27 (Miridiba castanea) 7 9, (Gametis jucunda) 4 7. 4 2 214. 101 (37 ). 5 4 45 113 2. 112 2. 101 56,457, 5,358. 2,000 4,000 7 8. 13 10. : Leach (1815), Mulsant and Rey (1842, 1871), Kolbe (1886), Heyden (1887), Bates (1888), Kraatz (1888), Reitter (1895), Lewis (1895, 1896), Arrow (1913), Nijiima and Kinoshita (1923, 1927), Murayama (1935b, 1937a), Cho (1969), Stebnicka (1980), Kalinina (1989), Kim (1992, 1993, 2000, 2001); : Murayama (1931), Medvedev (1952), Zhang (1984), Klausnitzer and Krell (1996). Family Cetoniidae Leach, 1815 Cetonida Leach, 1815: 99.,., ( )..,,., (Tribe Cremastochilini, : ). : Cetonia Fabricius, 1775: 42. : 12 ( 3 ). : 515 ( 14 ), 107 ( 9 ), 43 ( 3 ), 33 ( 2 ). : 4,000, 500 (Gorodinski, 2009) ( 18 ). :,. : Kolbe (1886) 1 7, Heyden (1887) 4. Kraatz (1888), 3. Cetonia metallica Protaetia 1. Bates (1888) 2 Glycyphana

28 kuperi Gametis jucunda, Niijima and Kinoshita (1923) 3 1. Murayama (1935), Cho (1969), Stebnicka (1980), Kim (1980), Kim and Chang (1982) 2 4, Cho (1934, 1957), Sawada (1937), Cho et al. (1967), Hyun and Woo (1969), Gu (1973) 1., 3 Cetonia (Kim, 1994) 1 2. (Valginae), (Trichiinae), (Cetoniinae) 3 30 70 46. 15 18. : Kolbe (1886), Heyden (1887), Lewis (1887), Bates (1888), Kraatz (1888), Reitter (1898), Nijiima and Kinoshita (1923, 1927), Arrow (1941), Mikšić (1976, 1977, 1979, 1982, 1987), Krikken (1977, 1984), Kim (1980, 1994), Stebnicka (1980), Kalinina (1989), Loebl and Smetana (2006), Smetana and Andrew (2006); internet Gorodinsk; : Murayama (1931), Medvedev (1952), Zhang (1984), Kim et al. (2003). 1., Cetoniinae, 2 2., 1 3. Trichiinae, 3 7.,,. 15 mm Valginae Subfamily Valginae Mulsant, 1842 Vulguaires Mulsant, 1842: 519. Valgaires Mulsant et Rey, 1871: 720. Valgini Seidlitz, 1875. 4 10 mm,.., 3 7. 5 5 2 2. : Valgus Scriba, 1790: 519.

: : 29 : 12 (Gorodinski, 2009) ( 2 ). : 111 (Gorodinski, 2009) ( 2 ). :,. : Sawada (1939, 1941a, 1944), Medvedev (1964), Krikken (1978a), Kobayashi (1983), Kim (2001a, b); : Medvedev (1952), Klausnitzer and Krell (1996). 1.. 3 7. gen. Nipponovalgus, angusticollis. 5, 1 3. gen. Valgus, koreanus Genus Valgus Scriba, 1790: 66. : Scarabaeus hemipterus Linné, 1758: 351. : 20 ( 1 ). : ( ), (3 ), (1 ). : Sawada (1944), Kim (2001b). 1. Valgus koreanus Sawada, 1944: 12 ( 1-1, 10-1, 17-1) Valgus koreanus Sawada, Trans. Kansai Ent. Soc., 14(1): 12 13, 1944; Kim, 2001b: 143. Valgus hemipterus Linné, Syst. Nat. ed. 10, I: 351, 1758 (Scarabaeus)( ): Murayama, 1937a: 36; Sawada, 1939: 82; Miwa and Chûjô, 1939: 78; Cho, 1957: 301; KZS, 1968: 138 ( ); Hyun and Woo, 1969: 194; Nam and Kim, 1983: 129 (Kim, 2001a: ). Chromovalgus fumosus Lewis, Ann. Mag. Nat. Hist., (5)19: 201, 1887 (Valgus)( ): Murayama, 1937a: 36; Sawada, 1939: 90; Miwa and Chûjô, 1939: 78; Cho, 1957: 301; KZS, 1968: 138 (

30 ); Won and Choi, 1968: 379 (Kim, 2001a: ). Valgus fumosus: Sawada, 1941: 3. Charitovalgus fumosus: Kobayashi, 1983: 2. Valgus pictus fumosus: Sawada, 1941: 3. Valgus sp. 1: Kim, 2001b: 130. :.,.,.,. 1.,. 5.,.,. Ad:,, ( )..,.,. BL: 7 8 mm ( ), 8 9 mm ( ), BW ( ): 3.6 4.2 mm ( ), 3.3 4.3 mm ( ). :. :,. : SK-GW:,,, GS:,,,,,, CBN:,, JBN:,,. (25ex): [SWU] : 4 2 ( : 22.iii.1990, Kim); 1 ( : 24.v.1993, Ku); 1 ( : 26.v.1984, Chang). (?): 1 3 ( : 14.v.1983, Chang); 1 ( : 14.v.1981, Chang); 1 ( : 14.v.1981, Chang); 1 ( : 9.v.1990, Kim); 1 ( : 9.v.1990, Kim); 1 (14.v.1967, Park); 1 (18.v.1997, Lee). : 1 ( : 19.v.1990, Kim). : 2 ( : 23.v.1997, Kim). : 2 ( : 21.v.1992, Kim and Oh); 1 ( : 26.v.1994, Kim K.R). : 1 ( : 26.v.1994, Hong). Genus Nipponovalgus Sawada, 1941: 4. Sawada (1941) 7. Dasyvalgus. : Valgus angusticollis Waterhouse, 1875: 115.

: : 31 : 2 ( 1 ). :,. : Sawada (1941). 2. Nipponovalgus angusticollis (Waterhouse), 1875: 115 ( 1-2, 10-2, 17-2) Valgus angusticollis Waterhouse, Trans. Ent. Soc. Lond.: 115, 1875: Heyden, 1887: 256. Dasyvalgus angusticollis: Niijima and Kinoshita, 1923: 199; Okamoto, 1924: 181; Murayama, 1935b: 6; 1937a: 36; Haku, 1937: 121; 1937: 74; Mochizuki and Tsunekawa, 1937: 91; Sawada, 1939: 87; Miwa and Chûjô, 1939: 78; Cho, 1957: 302; 1963: 221 ( ); KZS, 1968: 138; Cho et al., 1968: 265; Stebnicka, 1980: 287. Nipponovalgus angusticollis: Sawada, 1941: 4; Kim and Kim, 1975b: 256; Kim and Nam, 1982b: 157; 1983: 276; Kim, 1978: 373; Kim, 1993: 66; 2000a: 141; 2001: 129; 2005: 80; 2006: 37. : ( ).,.., 2. 7 5 7. 1,.. 1 2,3. Ad:. 1/2, ( ).,,. BL: 4 7 mm. BW: 2 3 mm. :. :,. :,,. : NK-, SK- :, GS:,,, -G, -G,,,,,,, CBN:,,,,,, GB:,,,,, JBN:,,,,,,,, JJ:. (187ex): [SWU] : 1ex ( : 14.vi.1997). : 22ex (, : v x.1971 1996); 1ex ( : 14.v.2006); 23ex ( : iv vi.1941 1997); 6ex ( : v vi.1989 1997). : 10ex (,,, : v.1984 1997). : 7ex (,,,, : v vi.1978 1990); 8ex

32 ( : 29.v.1999). : 14ex (,,,,,, : iv v.1987 1995). [NAAS] : 12ex (,, : v.1975, 1986). : 2ex ( : 12.v.1997). [SNU] : 22ex ( : iv vi.1990 1992). [KU] : 1ex ( : 8.vi.1974). : 1ex [Seiryori (= ): 4.v.1934]; 2ex [Mt. Namzan (= ): 21.iv.1935]; 50ex (,, : iv ix.1960 1989). : 1ex ( : 26.v.1974). : 1ex ( : 5.vi.1989). : 1ex ( : 23.v.1988). [HSU] : 1ex ( : 20.iv.1997). [HMNH-NK] 1ex ( : 26.v.1985). : 176, 10.2. 6.11. (iv- 13, v- 136, vi- 22, ix- 2, x- 3ex). Subfamily Trichiinae Fleming, 1821: 21. Trichiidae Fleming, 1821: 50.,. 6 65 mm.,.. Hairy Flower Chafer Bee-like Flower Scarab. 3 9 (Kim, 2001b) 3 3. : Trichius Fabricius, 1775: 40. : 43 ( 8 )( 3 ). : 82 (Gorodinski, 2009) ( 3 ). :. : Krikken (1978), Kim (2001b), Gusakov (2002), Tauzin (2006); : Klausnitzer and Krell (1996). 1. 3, 2.., gen. Osmoderma., 2 opicum 2, 1 2

: : 33 2. 1,,. gen. Trichius 12 mm., succinctus 1,.., gen. Gnorimus,. subopacus Genus Osmoderma Le Peletier de Saint-Fargeau and Serville, 1828: 702. Gymnodi Kirby, 1827. Gymnodus Kirby, 1837... 3 2,. : Scarabaeus eremitus Scopoli, 1763: 7 (MacLeay, 1838: 16 ). : 15, 10 ( 1 ). :. : Audisio et al. (2007). 3. Osmoderma opicum Lewis, 1887: 49 ( 1-3, 10-3) Osmoderma opicum Lewis, Wien. Ent. Zeit., VI: 49, 1887: Cho, 1969: 688 ( ); Kim, 1978: 379; ESK and KSAE, 1994: 153; Kim, 1998c: 358; 2001: 132. Osmodera apica: Nakatomi, 1934: 655. :.,.,

34 2, 2.. 3, 2.,.,.. BL: 22 35 mm, BW: 14 17 mm. :,. :. : SK-GW:,,. (5ex): [SWU] : 3 ( : 25 29.vii.2000, H.N. Seok); 1 ( : 9.viii.2000, J.R. Cha). [KU] : 1 ( :?). :,. Osmoderma = 1. Osmoderma barnabita Motschulsky, 1845 Osmoderma barnabita Motschulsky, 1845. Bull. Moscou., 18, I: 58, 1845; Stebnicka, 1980: 207 ( ); ESK and KSAE, 1994: 153 ( ), O. opicum. 2. Osmoderma caeleste Gusakov, 2002 Gymnodus caelestis Eidos, Kaluga: 39, 2002, (,, ), Tauzin (2006) O. barnabitum var. castaneum Tauzin, 1994 (, infra-subspecific name). Genus Gnorimus Le Peletier de Saint-Fargeau and Serville, 1828: 702. 1,.,. 1.

: : 35 : Scarabaeus nobilis Linné, 1758. Syst. Nat., ed. 10, 1: 358. (Blanchar, 1845 ). : 9 (internet: Gorodinsk) ( 1 ). :,,. : Tauzin (2004), Krell et al. (2006). 4. Gnorimus subopacus Motschulsky, 1860: 134 ( 1-4, 10-4, 17-4) Gnorimus subopacus Motschulsky, Schrensk s Reisen Amurl.: 134, 1860: Niijima and Kinoshita, 1923: 245; Cho, 1934: 75 (subofascus); 1957: 301; Murayama, 1935b: 6; 1937b: 36; Mochizuki and Tsunekawa, 1937: 91; Doi, 1938: 97; Miwa and Chûjô, 1939: 76; KZS, 1968: 138; Stebnicka, 1980: 289; Kim, 1982a: 157; Kim, 1993: 66; 2001b: 132; ESK and KSAE, 1994: 153 ( ). Gnorimus subcostatus subopacus: Heyden, 1887: 255. : (KZS, 1968: 138).,.... 2.,.. Ad:. 1.8. ( ). BL: 15 22 mm, BW: 7 8 mm. : Zhang, 1984: 32. :,,,. :,,,. : NK- HB:,,,, HN:,,, GW:, SK-GW:,,,,, GS:,,,,,,,, CB:, CN:, GB:,,, JJ:. (179ex): [SWU] : 2ex ( : 8.vi.1985); 1ex ( : 23.vii.1986). : 66ex (,,,, : iv vi.1935 1999). : 1ex ( : 20.vi.1992); 1ex ( : 18.v.2001). : 1ex (2.v.1992). [NAAS] : 1ex ( : 9.vii.1923). : 1ex ( : 3.vii.1931). [KU] : 1ex ( : 4.viii.1933). : 8ex (,, : v vi.1970 1979). : 77ex (,,,,,, : iv vii.1932 1993). : 1ex ( : 18.v.2001). : 2ex ( : 26.v.1974). : 5ex (,, : vi.1974 1996).?: 2ex (Koshcu Heihoku: 15.viii.1936); 1ex (Sounando cosho: 5.viii.1937). [HMNH-NK] : 6ex ( : 2 5.vi.1991). : 2ex ( : 27.v.1985).

36 : 171, 4.28. 11.10. (iv- 2, v- 100, vi- 58, vii- 5, viii- 4, ix- 1ex). Genus Lasiotrichius Reitter, 1899: 101... fasciatus Lasiotrichius. succinctus Lasiotrichius. : Scarabaeus succinctus Linné, Pallas, 1781. : 19 ( 1 ). :. 5. Lasiotrichius succinctus (Pallas), 1781: 18 ( 1-5, 10-5, 17-5) Scarabaeus succinctus Pallas, Icon. Ins.: 18, t. A, f. 19, 1781. Trichius succinctus Pallas: Kolbe, 1886: 196; Heyden, 1887: 256; Bates, 1888: 378; Niijima and Kinoshita, 1923: 194; Okamoto, 1924: 181; Kim, 1960: 27; Cho et al., 1967: 198; Hyun and Woo, 1969: 194; Kim et al., 1971b: 160; 1978: 134; 1981: 125; 1984: 328; 1985: 105; Kim, 1978: 378; Kim et al., 1982a: 146; 1987b: 506; 1994: 111; 1999: 2002: 120; 2003: 127; 2004: 117; 2005: 80; 2006: 37; 130; Kim, 1981: 345; 1993: 66; 2000a: 141; 2001b: 132; Nam and Kim, 1983: 129; Yoon et al., 1978: 83; 1979a: 150; 1979b: 150; 1990b: 111; Park and Cho, 1986: 128; Park et al., 1987: 673; Lee and Ko, 1988: 211; Lee et al., 1994: 147; ESK and KSAE, 1994: 153. Trichius succinctus Motschulsky: Kamijo, 1932: 21. Lasiotrichius succinctus Motschulsky: Niijima and Kinoshita, 1927: 75; Cho, 1934: 75; 1947: 65 ( ); Mochizuki, 1935: 32; 1936: 211; Mori and Cho, 1940: 13. Lasiotrichius succinctus Pallas: Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 76; Doi, 1938: 97; Nagaoka, 1938: 25; Cho, 1957: 301; 1963: 221; 1969: 688; Cho et al., 1968: 265; KZS, 1968: 138; Kim et al., 1972a: 84; 1974a: 109; 1974b: 230; 1975: 253; 1976: 102; 1977: 131; 1982a: 156; 1982b: 277; Stebnicka, 1980: 288; Kim and Chang, 1987a: 224; Lee et al., 1994: 147. Trichius abdominalis Erichson: Kim, 1956: 343; 1958: 98; Ku, 1963: 28. : (Kim, 1960; Cho et al., 1967), ( ) (Cho, 1969), (Kim et al., 1975 10 ).

: : 37, 3.,.,.,.,. Ad:,.,. BL: 9 12 mm, BW: 5 7 mm. : Zhang, 1984: 32 (L. succinctus). :,,,. :,,,,. : NK-HB:,,,, HN:,,,,,, PB:,,, PN:,, GW:, WH:, GG:, SK- (16 ) 130 170 3,000. (2,183ex): [SWU] NK: 1ex [Ketzanchin (= ): 6.v.1938]. : 180ex (,,,,,,,,,,,,,, : iv viii.1968 2001). : 225ex (,,, : v x.1974 2001). : 90ex (,,,,,,,,, : v viii.1986 2003); 1ex ( : 7.vi.2008); 7ex ( : 7.vi, 5.vii.2008). : 232ex (,,,,,,,,,,,,,,, : v x.1977 2000); 2ex ( : 7.vii.2008); 2ex ( : 6.vi.2008); 2ex ( : 8.vi.2008). : 56ex (,,,,, : v viii.1987 1997). : 13ex (vii, viii.1983, 1988); 1ex ( : 15.vii.2007). [NAAS] NK: 3ex [Sharei (= ), Shakuoji (= ): vi viii.1922, 1923]; 1ex [Futenpo (= ): 3.vii.1931]. : 3ex [Gesseiji (= ): vii ix.1923]; 6ex (,, : vi ix.1961 1997). : 14ex ( : v vi.1957 1991). : 2ex (, : vii, viii.1987, 1995). : 20ex (, : vii ix.1988 1996). : 1ex [Hakuyosan (= ): vi.1922]; 2ex (, : vi, vii.1963, 1993); : 1ex [Yotouri ( ): 8.vi.1931]. [SNU] : 7ex (,,, : vii ix.1924 1997). : 203ex (,,,,,,,,,, : iv ix.1969 1997). : 31ex (,, : vi xi.1983 1995). : 3ex (,, : vi, viii.1994 1997). : 86ex (,,,, : v ix.1964 1997). : 2ex ( : vii.1971).?: 6ex ( : vii viii.1993). [KU] NK- HN: 6ex [, Keizanchin Kannan (= ): v viii.1933]; 2ex [Kazando Cocho-c (?): vii, viii.1937]; 2ex ( : 16.vii.1930). : 236ex (,,,,,,, : v viii.1967 1994). : 377ex (,,,,,, : iv x.1932 1996). : 15ex (,, : v viii.1957 1991). : 155ex (,,,,,,, : vi viii.1960 1996). : 76ex (,,,,, : iv viii.1970 1993). : 6ex (vii viii.1955 1974). [HNHM-NK] : 25ex ( : 13 18.vii.1982); 3ex ( : 8.vi.1991). : 3ex ( : 10.vi.1970); 6ex (16.vi.1985). : 1ex ( : 24.v.1970). : 3ex ( : 29.v.1970); 1ex (1.vi.1970); 6ex (6.viii.1975); 1ex (10.vii.1977); 13ex (18 21.vi.1985); 12ex (12.vi.1991); 1ex (12.vii.1991). : 5ex ( : 6 8.vi.1970). : 2,092, 4.14. 11.23. (iv- 10, v- 109, vi- 989, vii- 488, viii- 386, ix- 29, x,xi- 6ex).

38 (= ) 1. Paratrichius septemdecimguttatus (Snellen and Vollenhoven), 1864 Trichius septemdecimguttatus Snellen and Vollenhoven, Tijdschr. Ent. Nederl.: 159, 1864 (China, Hubei). Gnorimus septemdecimguttatus (Snellen and Vollenhoven): Sawada, 1937a: 33 ( ); Miwa and Chûjô, 1936: 76; Cho, 1957: 301; ESK and KSAE, 1994: 153 ( ). Subfamily Cetoniinae Leach, 1815: 99. Cetonida Leach, 1815. ( ), ( ),.,.. 61 4 9 13. : Cetonia Fabricius, 1775: 42. : 10 [ 7 (Gorodinski, 2009)] ( 4 ). : 100 [ 72 (Gorodinski, 2009)] ( 9 ). : 4,000 [ 500 (Gorodinski, 2009)] ( 13 ). :. : Kolbe (1886), Heyden (1887), Lewis (1887), Bates (1888), Kraatz (1888), Reitter (1898), Nijiima and Kinoshita (1923, 1927), Murayama (1935, 1937), Arrow (1941), Mikšić (1976, 1977, 1979, 1982, 1987), Krikken (1977, 1984), Kalinina (1989), Kim (1994b), Smetana and Andrew (2006); internet: Gorodinsk; : Murayama (1931), Medvedev (1952), Zhang (1984), Kim et al. (2003). 1.. tribe Cremastochilini,...

: : 39 gen. Clinterocera 12 mm,. 1 obsoleta ( )., 2 2..,. tribe Phaedimini,.. gen. Dicranocephalus, adamsi, 3 3.,. tribe Coryphocerina 4,, tribe Cetoniini 4..,. Ad: gen. Rhomborhina. 1,. polita. gen. Pseudotorynorhina., 1. japonica Tribe Phaedimini Schoch, 1894: 169. Phaedimini Schoch, 1894: 169. 5 30 mm,.. 5 27 (Mikšić, 1977), 7 Dicranocephalus 1.

40 : Phaedimus Westwood, 1841a: 5. : 5 (Mikšić, 1977) ( 1 ). : 27 (Mikšić, 1977) ( 1 ). :,. : Murayama (1929a, 1930a), Mikšić (1977). Genus Dicranocephalus Hope, 1831: 24. : Dicranocephalus wallichi Hope, 1831. : 7 (Mikšić, 1977) ( 1). :. 6. Dicronocephalus adamsi Pascoe, 1863: 25 ( 1-6, 10-6, 17-6) Dicranocephalus adamsi Pascoe, Journ. Ent., II: 25, 1863: Niijima and Kinoshita, 1923: 145; 1927: 66; Murayama, 1929: 43; 1930: 57; 1937c: 93; Tanaka, 1935: 163; Cho, 1936: 26; 1956: 77 ( ); 1957: 302; 1969: 690 ( ); Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 79; Kobayashi, 1942: 500; KSZ, 1968: 138; Kim et al., 1974b: 230; 1975: 253; 1982a: 157; 1982b: 277; 1985: 105; Mikšić, 1977: 171; Kim, 1978: 380; Stebnicka, 1980: 281; ESK and KSAE, 1994: 154; Lie et al., 1997: 87; Kim, 1998c: 358; 2001b: 138.,.,,,..,.,. 1.5,..,. Ad: 1.5.

: : 41,, ( ).. BL: 27 35 mm ( ), 21 27 mm ( ), BW: 10 13 mm (, ). :,,,. :,. : NK-WH:,, GG:, SK-GW:,,,,,, GS:,,,,,,,,,,,, (6 ),,,,,,,,, CBN:,,,, GBN:,,,, JBN:,,,. (307ex): [SWU] : 1ex ( : 26.v.1984). : 57ex (,,,, : v viii.1978 1999). : 2ex (, : vii.1991, v.1988). : 1ex ( : 21.ix.1989). : 2ex ( : 21.v.1992). [NAAS] : 2ex (, : v.1982, 1989). [SNU] : 1ex ( : 26.v.1985). : 21ex (,, : v vi.1968 1992). : 1ex ( : 24.vi.1995). [KU] : 1ex ( : 5.vi.1967). : 204ex (,,, : iv x.1935 1990). : 3ex ( : vi.1997). : 3ex (, : v vii.1985, 1986). 6ex ( : v.1991). : 294, 4.21. 10.16. (iv- 3, v- 139, vi- 139, vii- 6, viii- 2, ix- 1, x- 4ex). Tribe Coryphocerina Burmeister, 1842: 215. Coryphoceridae Burmeister, 1842: 215...,. 19 Rhomborrhina 2 Pseudotorynorrhina 1. : Coryphocera Burmeister, 1842: 220. : 24 (Mikšić, 1977) ( 2 ). : 92 (Mikšić, 1977) ( 2 ). :. : Mikšić (1977).

42 Genus Rhomborrhina Hope, 1837: 120. : Goliathus heros Gory and Percheron, 1833. : 19 (Gorodinski, 2009) ( 1 ). :. 7. Rhomborrhina polita Waterhouse, 1875: 113 ( 1-7, 10-7, 17-7) Rhomborrhina polita Waterhouse, Trans. Ent. Soc. Lond., 1875: 113, 1875: Kim, 1960: 27; Cho et al., 1967: 197; Cho, 1969 ( ): 693; Kim, 1978: 381; Kim and Nam, 1984: 27; Yoon and Nam, 1985: 160; ESK and KSAE, 1994: 154; Kim, 2001b: 139. : ( ) (Cho et al., 1967 3). : (Yokohama)..,..,,,.,. 1.,.. BL: 27 35 mm, BW: 14 17 mm. :,. :. : SK-GW:, CN:, GN:, JJ:,. (3ex): [SWU] : 1 ( : 19.viii.1935). : 1ex ( : 7.viii.1999, S.Y. Kim). [NAAS] : 1 ( : 9.viii.1934).

: : 43 Genus Pseudotorynorrhina Mikšić, 1974: 64. : Rhomborhina japonica Hope, 1841. : 2 ( 1 ). :. : Mikšić (1976, 1977). 8. Pseudotorynorrhina japonica (Hope), 1841: 64 ( 2-8, 10-8, 17-8) Rhomborrhina japonica Hope, Trans. Ent. Soc. London, III: 64, 1841: Kolbe, 1886: 194; Niijima and Kinoshita, 1923: 156; Okamoto, 1924: 179; Doi, 1935: 3; Morita, 1936: 860; Haku, 1937: 696; Kamijo, 1936: 86; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô 1939: 81; Takahashi, 1941: 235; Kim, 1956: 339; Cho, 1957: 302; 1963: 220 ( ); 1969: 692; Cho et al., 1968: 265 (Phomborrhina); KZS, 1968: 138; Hyun and Woo, 1969: 194; Gu, 1973: 82; Kim and Kim, 1974a: 109; 1975: 254; Kim and Nam, 1982b: 277; Kim, 1978: 381; Yoon et al., 1979a: 83; 1985: 160; 1989: 140; Stebnicka, 1980: 281; Kim et al., 1982a: 145; 1982b: 173; 1984: 169; 1987b: 506; 1989: 177; Kim, 1983: 166; Lee and Ko, 1988: 211. Pseudotorynorrhina japonica: Mikšić, 1976: 261; Kim and Yoo, 1987: 224; Kim et al., 2001b: 139; 2002: 120; ESK and KSAE, 1994: 154. Rhomborrhina japonica coreana Ruter, Bull. Soc. Ent. Fr. 1965: 196. Rhomborrhina nigra Saunders, Trans. Ent. Soc. Lond., II: 29, 1852 ( ): Cho 1969: 692. Rhomborrhina unicolor Motschulsky, Etud. Ent., X: 8, 1861 ( ): Murayama, 1937: 36; Takahashi 1941: 235; KZS, 1968: 138. : (Gu, 1973), (KSPP, 1972); nigra= (Cho 1969); unicolor= (KZS, 1968). :.,.,,,.,.,. 3,,.. 1.

44.. Ad:.,.. ( ). BL: 25 33 mm, BW: 12 17 mm. : Kamijo, 1936: 86. :,,. :,,. : NK-, SK- (6 ) 60 80 500. (310ex): [SWU] : 1ex ( : 14.vii.1997). : 42ex (,,,,,, : v x.1971 1992). : 5ex (,, : vii viii.1984 1997). : 4ex (,, : vii, viii.1990 2000). : 24ex (,,,,, : vii viii.1980 1996); 5ex ( : 15, 16.vii.2003). : 163ex ( : vi viii.1980 2002). [NAAS] : 1ex ( : 16.viii.1989). : 2ex (, : ix.1992, vi.1984). : 4ex (, : vii, viii.1991, 1996). : 1ex [Hakuyosan (= ): 25.vi.1922]; 6ex (,,, : vii viii.1959 1993). : 2ex (vii, viii.1993, 1995). [KU] : 4ex (,, : vii x.1966 1982). : 5ex (, : vii.1957, 1982). : 4ex ( : vii.1978). : 31ex (,,,,, : vii, viii.1955 1990).?: 5ex (Zokani: 14.viii.1930). [HSU] : 1ex ( : 17.vi.1997). : 182, 5.4. 10.24. (v- 2, vi- 6, vii- 115, viii- 51, ix- 4, x- 4ex). Tribe Cetoniini Leach, 1815: 99. Cetonida Leach, 1815., Kolbe (1886) Cetonia seulensis 8 3, Gu (1973) C. roelofsi. 15 48 5 9. : Cetonia Fabricius, 1775: 42. : 110 [ 30 (Mikšić 1982, 1986; 16: Gorodinski, 2009)] ( 5). : 1,000 [ 450 (Mikšić 1982, 1986), 200 (Gorodinski, 2009)] ( 9). :.,,. : Kolbe (1886), Heyden (1887), Lewis (1887), Bates (1888), Kraatz (1888), Reitter (1898),

: : 45 Niijima and Kinoshita (1923, 1927), Murayama (1935, 1937), Kato (1937), Reitter (1898), Mikšić (1982, 1987), Kalinina (1989), Kim (2000b, 2001b); : Kim et al. (2003). 1., 2,.,. gen. Anthracophora,. 9 1, 3, 5, 6 rusticola 2.,. 3. 4 3.,.,. Ad: gen. Glycyphana,. fulvistemma.,. Ad: gen. Protaetia 5 4. 1.,.., gen. Gametis. 1 jucunda.,.. gen. Cetonia.., *, pilifera pilifera *, pilifera magnifica

46 Genus Cetonia Fabricius, 1775: 42....... 8 3 Protaetia, 1 2. : Scarabaeus auratus Linné, 1758. : 29 (Gorodinski, 2009) ( 1 2 ). :. : Mikšić (1982), Kim (1994b, 2001b). 9. Cetonia pilifera (Motschulsky), 1860: 15 ( 1-9, 10-9, 17-9/1-9a, 10-9a, 17-9a) Glycyphana pilifera Motschulsky, Etude. Entom., IX: 15, 1860; Bates, 1888: 377. Cetonia pilifera: Okamoto, 1924: 179; Miwa and Chûjô, 1939: 86; Kim, 1958: 92; 1960: 27; Cho, 1963: 220 ( ); Cho et al., 1967: 198; 1968: 265; Gu, 1973: 82; Stebnicka, 1980: 209; Mikšić, 1982: 221; Kim and Nam, 1984: 328. Eucetonia pilifera: ESK and KSAE, 1994: 154; Park and Park, 1987: 673. Cetonia (Eucetonia) pilifera pilifera (Motschulsky): Kim, 1994b: 317 ( ); 2001b: 141. Cetonia (Eucetonia) pilifera magnifica Ballion: Kim, 1994b: 317 ( ); 2001b: 141; 2002: 120. Cetonia aurata var. pilifera: Heyden, 1887: 255. Cetonia magnifica Ballion, Bull. Soc. Nat. Moscou, XLIII: 348, 1870: Cho, 1934: 75; Murayama, 1935b: 5; 1937a: 36; Mochizuki, 1935: 32; 1936: 211; Mochizuki and Tsunekawa, 1937: 90; Doi, 1938: 97; Miwa and Chûjô, 1939: 86; Stebnicka, 1980: 282; Mikšić, 1982: 223; Kalinina, 1989: 433; Kim, 1993: 66. Glycyphana cupreola Kraatz, Deut. Entom. Zeit., 23: 243, 1879; Bates, 1888: 376. Cetonia aurata var. cupreola: Heyden, 1887: 254. Cetonia viridiopaca (Motschulsky), Schrenk s Reis. Amurl.: 134, 1860 (Glycyphana): Stebnicka, 1980: 209; Mikšić, 1982: 223; Lie et al., 1997: 87. Eucetonia viridiopaca: ESK and KSAE, 1994: 154. Cetonia aurata var. viridiopaca: Heyden, 1887: 254. Cetonia roelofsi Harold, Bull. Soc. Ent. Belg. p. 8, 1880 ( ): Gu, 1973: 82; Stebnicka, 1980: 209; Kim

: : 47 and Nam, 1984: 328; Yoon and Nam, 1986: 160; Kalinina, 1989: 433. : pilifera= (Kim, 1958, 1960), (Gu, 1973); roelofsi= (Gu, 1973; ESK and KSAE, 1994), (KSPP, 1972). : Vladivostok (?)...,.. 3. 2,.. (magnifica),. (pilifera),. Ad: 1.2.,. 1.7. BL: 16 20 mm, BW: 8 10 mm. : Medvedev, 1952: 178 (C. magnifica). :,,,. :,,,. : NK-HB,,,,,,, HN,,,,,,,,,, PB,, PY,,, WN,, GG, SK- (2ex) 45 60 600. (562ex): pilifera=[swu] : 12ex (,, : 4.v.1978). magnifica=[swu] : 6ex (,,, : v viii.1958 1990). : 81ex (,,,,,, : iv ix.1957 1998); 1ex ( : 3.vii.2007); 1ex (4.viii.1982). : 8ex (,, : v viii.1986 1989). : 5ex (, : vii.1983, 1984). : 7ex (v.1978 1990); 2ex (7, 10.vii.2002). [NAAS] NK- : 5ex [Sharei (= ), Shakuoji (= ), Kankakei (= ), Futenpo ( ): vi vii.1923, 1924, 1931]. : 1ex ( : 10.iv.1923); 4ex (, : v vii.1975, 1976). : 3ex ( : 21.vi.1973). : 4ex (28.vi.1922).?: 1ex (Koryosan: v.1924). [SNU] : 20ex (,, : v vi.1969 1992). : 5ex (v.1995). [KU] NK-11ex [ -Keizanchin Kannan (= ), Fusenrei (= ), -Gosui (= ),?-Sounando Cosho, Kousho Heihoku: vii viii.1931 1937]; : 1ex ( : 17.vi.1967); : 304ex (,,,,,,,,, : iv xi.1932 1985)., : 12ex (,, : v viii.1972, 1974, 1983, 1985). : 1ex ( : vii.1978); : 3ex ( : vii.1979). : 2ex (v.1989, 1990). [HMNH-NK] 31ex [ (1,150 m) YG: 22, 26.vii.1975]; 27ex [ (, 1,100 m) YG: 27.vii.1975]. : 2ex ( : 2.vi.1991). : 1ex ( : 30.v.1970). : 1ex ( : 30.vii.1982). : 478, 4.12. 11. (iv- 12, v- 147, vi- 138, vii- 149, viii- 26, ix- 4, xi- 2ex).

48 Genus Protaetia Burmeister, 1842: 22, 827. Protaetia Burmeister, 1842.,...... Kolbe (1886) Cetonia seulensis (=Protaetia brevitarsis seulensis) 14 5. : Cetonia spectabilis Schaum, 1841. Handb. Ent. III: 473. : 250 (Ratecliffe, 2002) [ 123 (Gorodinski, 2009)] ( 5 ). :,, ( 1 : Ratecliffe, 2002). : Niijima and Kinoshita (1923, 1927), Murayama (1935, 1937), Kato (1937), Mikšić (1987), Kim (2000b, 2001b); : Klausniter and Krell (1996), Kim et al. (2003). 1. 2 ( ), 2. 2 1. 3 2.. Ad: (subg. Calopotosia) orientalis submarmorea. Ad: ( ), (subg. Chrysopotosia) mandschuriensis 3.. (subg. Liocola) 4, (subg. Potosia).. Ad: famelica scheini 4. brevitarsis seulensis, lugubris

: : 49 10. Protaetia lugubris (Herbst), 1786: 157 ( 1-10, 11-10, 17-10) Cetonia lugubris Herbst, in Füessly, Arch. Insectengesch. 7 8: 157, 1786. Protaetia lugubris: Mikšić, 1987: 496; ESK and KSAE, 1994: 154 ( ); Kim, 2000b: 213; 2001b: 144. Potosia (Liocola) lugubris orientalis Medvedev, Faun. SSSR, Col. X, 5: 185, 1964: Stebnicka, 1980: 209. Nectosia (Liocola) lugubris orientalis: Kalinina, 1989: 434.,..., 3..,.,. Ad:, 1.5..,. BL: 19 24 mm, BW: 12 14 mm. :,,,. :,,. : SK-GW:,, GS:,,,,,, GB:, GN:, JB:,,?: Sounando Cosho-c. (21ex): [SWU] : 1ex ( : 6.vi.1985); 1ex ( : 1.vii.1984). : 1ex ( : 24.v. 1995). [SNU] : 1ex ( : 2.v.1990); 1ex ( : 13.vi.1973); 1ex ( : 10.vi.?).?: 1ex (Sounando Cosho-c: 14.vii.1937). [KU] : 1ex ( : 22.v.1977); 1ex ( : 10.vi.1995). : 3ex ( : v ix. 1935 1981); 1ex ( : 5.vi.1979); 1ex ( : 22.vi.1975). : 5ex ( : viii.1956, 1979). : 1ex ( : 9.vi.1972); 1ex ( : 26.v.1985). 11. Protaetia brevitarsis seulensis (Kolbe), 1886: 194 ( 1-11, 10-11, 17-11) Cetonia brevitarsis Lewis, Ann. Mag. Nat. Hist. (5)IV: 463, 1879; Kraatz, 1888: 200. Cetonia seulensis Kolbe, Arch. f. naturg., 52: 194, pl. XI, f. 29, 1886: Heyden, 1887: 254; Bates, 1888: 37. Liocola brevitarsis: Okamoto, 1924: 179; Masaki, 1934: 402; 1936: 260; Doi, 1935: 3; Yoshino, 1935: 14; Kato, 1937: 619; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 87; Cho, 1955: 207;

50 1957: 301; 1963: 220; 1965: 197; 1969: 693; Kim, 1956: 9; 1958: 98; Ku, 1963: 27; KZS, 1968: 138; Cho et al., 1968: 265; Hyun and Woo, 1970: 80; Kim et al., 1972b: 197 ( ); 1974a: 109; 1982a: 157; 1982b: 277; Yoon and Nam, 1986: 160. Protaetia brevitarsis: Kim, 1978: 383; KSPP, 1972: 208; Shin, 1978: 144; Yoon and Nam, 1979: 150; Lee and Kwon, 1981: 154; Kim and Nam, 1984: 328; 1985: 105; Kim and Chang, 1982a: 145; 1984: 170; 1987b: 506; Kim, 1993: 66; Lee et al., 1994: 147. Potosia (Liocola) brevitarsis: Stebnicka, 1980: 283; Lie et al., 1997: 88. Nectosia (Liocola) brevitarsis: Kalinina, 1989: 434. Pachnotosia brevitarsis seulensis: Reitter, 1898: 65. Liocola brevitarsis seulensis: Niijima and Kinoshita, 1923: 165; Kamijo, 1932: 21; Morita, 1936: 861; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 87; Cho, 1957: 302 (Seoulensis); 1969: 694; Gu, 1973: 85. Protaetia brevitarsis seulensis: Kim and Lee, 1979: 84; Kim and Chang, 1983: 166; Kim et al., 1994: 111; Kim, 2000b: 213; 2001b: 144; 2005: 80. Liocola brevitarsis viridis Kraatz, Deut. Entom. Zeit.: 380, 1889: Kamijo, 1932: 21; Haku, 1937b: 696; Miwa and Chûjô, 1939: 87. Liocola speculifera Swarz, 1817 ( ): Murayama, 1935: 6; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 88; Cho, 1947: 65; 1957: 302; 1963: 219; Kim, 1956: 9; 1958: 98; Cho et al., 1967: 198; 1968: 265; KSZ, 1968: 138; KSPP, 1972: 208 (Protaetia); Kim and Nam, 1984: 328. Protaetia speculifera: KSPP, 1972: 208. Protaetia pryeri Janson, 1888 ( ): Kim and Chang, 1982a: 145; 1987a: 224. Protaetia cataphracta Arrow, 1913 ( ): Kim and Chang, 1982a: 146; 1987a: 224; Kim and Yoo, 1987b: 506. : ( ) (Cho, 1955 2), (Cho, 1965 3), (Cho, 1963; Cho et al., 1968), (KSPP, 1972); speculifera= (Cho, 1947), ( ) (Cho et al., 1967 2), (Cho, 1963 2), ( )(KSPP, 1972). : Cetonia brevitarsis-. Cetonia seulensis-.,.,.. 3, 3. 2,.., 7. Ad: 1.3. 2, ( ),. BL: 17 22 mm, BW: 12 15 mm. : Murayama, 1931: 75; Medvedev, 1952: 180 (L. brevitarsis); Zhang, 1984: 27 (P. brevitarsis); Kim et al., 2003: 232.

: : 51 :,, ( ). :,,,,. : NK-HN, PN,,,? Sunan and Sǒkam-Čŭsudži, SK- (6 ) 70 90 600. (472ex): [SWU] : 4ex (,, : vi viii.1974 1984). : 108ex (,,,,,,, : vi ix.1975 1996). : 5ex (,, : vii.1985 1994). : 51ex,,,,, : v viii.1978 1996). : 19ex (,,,,, : vi x.1981 1992). : 63ex (viii.1980 1994). [NAAS] : 1ex [Geseiji (= ), 15.ix.1923]. : 25ex (,, : v ix.1923 1985). : 1ex ( : 22.vii.1975). : 6ex (vii ix.1974 1988).?: 1ex (GW ainei: 22.viii.1922). [SNU] : 1ex ( : 19.vi.1992). : 32ex (,, : v vii.1962 1994). ; 3ex ( : v.1991). : 1ex ( : 5.vi.1987). : 13ex ( : vii viii.1970 1973). [KU] NK: 1ex (Sounando Cosho-c: 17.vii.1937). : 2ex ( : vii, ix.1978, 1979). : 1ex [Seiryori (= ): 4.viii.1933]; 62ex (,,, : v ix. 1931 1988). : 6ex (, : viii.1955, 1957). : 13ex (,,, : vi viii.1956 1990). : 12ex (,,,,, : vii viii.1971 1979). : 31ex (vii viii. 1931 1990); 1ex ( : 3.v.1955).?: 1ex (Zokani: 14.viii.1930). [HMNH-NK] : 4ex ( : 9.viii.1971); 1ex ( : 22.viii.1991). : 1ex ( : 6.viii.1975); 1ex (9.vii.1977); 1ex (18.vi.1985). : 447, 5.1. 9.29. (v- 4, vi- 56, vii- 183, viii- 179, ix- 15ex). : Kamijo, 1936: 85; Nakayama and Okamoto, 1940: 199; Saito, 1941: 12. 12. Protaetia famelica scheini (Mikšić), 1959: 29 ( 2-12, 10-12, 17-12) Cetonia famelica Janson, Cist. Ent. II: 539, 1879. Potosia (Potosia) famelica: Stebnicka, 1980: 284; Lie et al., 1997: 89. Protaetia famelica: Paik, 1984: 184. Nectosia (Potosia) famelica: Kalinina, 1989: 434. Potosia famelica scheini Mikšić, Acta Mus. Maced. sc. nat. VI: 29, 1959. Protaetia famelica scheini: Mikšić, 1987: 262; ESK and KSAE, 1994: 154; Kim, 2000b: 212; 2001b: 143. Potosia famelica amurensis: Niijima and Kinoshita, 1923: 176; Miwa and Chûjô, 1939: 89. Potosia amurensis Heyden, Deut. Entom. Zeit., 30: 285, 1886: Cho, 1957: 130; KSZ, 1968: 138 ( ). : (Paik, 1984). : Protaetia famelica-. Potosia famelica scheini-..,,

52..,.,. 3 3.,,. ( 7 ). Ad:, 5.. 1.2,,.. BL: 14 18 mm, BW: 8 10 mm. :,,,. :,,,. : NK-HN, PN,, SK-GW,, GS -G, -G, -G,,, CBN,, GB,, JN,, JJ. (97ex): [SWU] : 2ex (, : vi, vii.1981, 1984). : 54ex (,,, : v vii.1980 1987). : 4ex (, : v vii.1991 1995). : 2ex ( : vi.1990). : 1ex ( : vi.1991); 1ex ( : 23.vi.1987). [KU] NK- : 2ex [Keizanchin kannan (= ): 5.vi. 1933]. : 27ex (,,,, : v x.1956 1986). : 1ex ( : 14.vi.1996). : 2ex (, : v vii.1979, 1991). : 1ex (21.viii.1968). : 92, 5.6. 10.10. (v- 13, vi- 30, vii- 33, viii- 14, ix, x- 2ex). 13. Protaetia mandschuriensis (Schürhoff), 1933: 29 ( 2-13, 10-13, 17-13) Calopotosia mandschuriensis Schürhoff, Mitteil. Deutsch. Entom. Gesell., IV, 3: 29, 1933. Protaetia mandschuriensis: Kim, 2000b: 215; 2001b: 148; 2004: 117.,..,.. 3, 1.,. 7,. 7,. Ad: 2/3, 1/8.,,. BL: 22 28 mm, BW: 13 15 mm.

: : 53 : Kim et al., 203: 234. :,,. :,,. : NK-PY, SK-GW, GS,,,,,,,, CN,, JB,,, JN. (91ex): [SWU] NK-3ex ( : vii viii.1975 1982). : 23ex (,, : iv viii. 1977 1996). : 4ex ( : 16 18.vi.2003). : 2ex ( : vii.1987). : 2ex (, : vi.1991, 1995). [NAAS] : 1ex ( : 27.v.1984); 2ex [Kasan (= ): 1.vi.1922]. [SNU] : 2ex ( : 27.vi.1997). : 8ex (,,, : v viii.1970 1993). : 4ex ( : 4.v.1991). : 1ex ( : 30.vii.1994). [KU] : 4ex (, : vi.1967, 1988). : 34ex (,,, : v ix.1934 1991). : 1ex ( : 4.viii.1974). : 91, 4.5. 9.21. (iv- 1, v- 20, vi- 48, vii- 10, viii- 11, ix- 1ex). 14. Protaetia orientalis submarmorea (Burmeister), 1842: 460 ( 2-14, 11-14, 17-14) Cetonia orientalis Gory-Perchron, Mon. Ceton.: 193, tab. 34. fig. 6, 1833. Cetonia submarmorea Burmeister, Hand. Entom., III: 460, 1842: Bates, 1888: 377. Protaetia orientalis: Ku, 1963: 28; Hyun and Woo, 1969: 194; Kim, 1978: 385; Kim et al., 1979: 84; 1982a: 145; 1983a: 166; 1989: 177; 2000a: 141; Lee and Kwon, 1981: 154; Kim and Nam, 1982a: 157; 1985: 105; Park and Park, 1987: 673; Kim, 1993: 66; Lee et al., 1994: 147. Potosia (Calopotosia) submarmorea: Reitter, 1898: 67. Protaetia orientalis submarmorea: Paik, 1984: 184 ( ); ESK and KSAE, 1994: 154; Kim, 2000b: 214; 2001b: 146; 2002: 120; 2004: 117; 2005: 80. Potosia aerata Erichson, Nova Acta Halle, 16: 240, 1834: Niijima and Kinoshita, 1923: 172; Okamoto, 1924: 180; Doi, 1935: 3; Miwa and Chûjô, 1939: 88; Cho, 1957: 302; 1963: 220; 1969: 194; KSZ, 1968: 138; Cho et al., 1968: 265; Won and Choi, 1968: 379; Shin and Noh, 1970: 37; Kim and Kim, 1971a: 54; 1972a: 84; 1972b: 197; 1974a: 109, 1974b: 230; KSPP, 1972: 208; Stebnicka, 1980: 284; Yoon and Nam, 1986: 160 ( ); 1989: 140; Mikšić, 1987: 541 (Syn. Nov.). Protaetia ishigakia Fairmaire, 1898 ( ): Kim, 1981: 345; Kim and Chang, 1982b: 173. Protaetia lewisi Janson, 1888 ( ): Kim, 1981: 345; Kim and Chang, 1982a: 146. : orientalis= (Hyun and Woo, 1969), (Kim, 1978 3); aerata= (Cho, 1963; Cho et al., 1968), (KSZ, 1968 10), (KSPP, 1972).

54 : Cetonia orientalis-. Cetonia submarmorea-..,..,.. 3, 3.,.,, 7. 3,, 7. Submarmorea,. Ad: 2. ( ).,.,. BL: 16 25 mm, BW: 11 15 mm. : Kim et al., 2003: 235. :,,,,,,, ; -,,. :,,,,. : NK-PY, HN, GW, SK- 100 130 1,000. (745ex): [SWU] NK-1ex ( : 10.viii.1971). : 7ex (,,,,, : vii viii.1974 1992). : 164ex (,,,,,,,,,, : iv x.1957 1997). : 17ex (,,,,,,,, : v viii.1967 2003). : 100ex (,,,,,,,,,,,,,, : iv viii.1979 1997). : 86ex (,,,,,,,,,,, : vi viii.1958 1997); 5ex ( : 15, 16.vii.2003). : 130ex (vi viii.1974 1994); 1ex ( : 16.viii.2009). [NAAS] : 1ex [Onseiri (= ): 25.vi.1924]; 1ex ( : 30.v.1980). : 1ex ( : 10.vi.1982); 11ex ( : vi xi.1965 1992). : 2ex [Tsudoji (= ): 25.vii. 1922]; 2ex (, : vii, viii.1993, 1998). : 2ex ( : vi, vii.1973, 1975). : 5ex (vii viii. 1922 1988). [SNU] : 7ex (,, : vi.1988 1992). : 58ex (,,,,, : iv ix.1969 1997). : 1ex ( : 24.vi.1986). : 5ex (,, : vi x.1987 1995). : 13ex (,,,, : vi viii.1986 1997). : 1ex (17.vii.1995). [KU] : 7ex (,,,, : iv viii.1968 1995). : 45ex (,,,,,,, : iv x.1932 1990). : 4ex (,, : vi vii.1957 1979). : 10ex (,,, : vi vii.1978 1990). : 53ex (,,, : v viii.1955 1990). [HSU] : 4ex ( : v vi.1996, 1997). : 615, 4.18. 11.18. (iv- 1, v- 27, vi- 114, vii- 242, viii- 202, ix- 22, x, xi- 7ex).

: : 55 Genus Anthracophora Burmeister, 1842: 624....,. : Anthracophora rusticola Burmister, 1842, Handb. Ent., III: 624. subg. Anthracophora: Cetonia crusifera Olivier, 1789, Ent. I, nr. 6, p. 39, tab. 5, fig. 29. subg. Poecilophilides: Anthracophora rusticola Burmister, 1842, Handb. Ent., III: 624. : 9 (3 : Mikšić, 1982), 7 (Gorodinski, 2009) ( 1 ). :,. : Mikšić (1982), Kim (2001b). 15. Anthracophora rusticola Burmister, 1842: 624 ( 2-15, 11-15, 17-15) Anthracophora rusticola Burmister, Handb. Ent., III: 624, 1842: Kolbe, 1886: 196; Heyden, 1887: 255; Niijima and Kinoshita, 1923: 164; Okamoto, 1924: 179; Kamijo, 1932: 21; Doi, 1935: 3; Mochizuki and Tsunekawa, 1937: 90; Miwa and Chûjô, 1939: 91; Cho, 1957: 302; 1963: 220; KSZ, 1968: 138; Ku, 1963: 27; Cho et al., 1968: 265 (susticola); Won and Choi, 1968: 379; Hyun and Woo, 1969: 193; KSPP, 1972: 206; Kim and Kim, 1971: 160; Mikšić, 1982: 487; ESK and KSAE, 1994: 154; Kim and Lee, 1997a: 231; Kim and Kim, 1998: 170; Kim, 1993: 66; 1998: 354; 2000a: 141; 2001b: 149; 2006: 37. Poecillophilides rusticola: Cho, 1969: 695; Kim, 1978: 386; Yoon and Nam, 1979b: 150; Stebnicka, 1980: 284; Kim and Kim, 1974a: 109; Kim and Nam, 1977: 131; 1982a: 157; 1982b: 277; 1984: 329; 1985: 105; Lee and Kwon, 1981: 154; Kim and Chang, 1982a: 146; 1987b: 506; Nam and Kim, 1983: 129; Lie et al., 1997: 89. : (Hyun and Woo, 1969; KSPP, 1972). : ( )...,.,., 9 1, 3, 5, 6. 7, 30. 7

56,. Ad:.,.. BL: 16 22 mm, BW: 8 11 mm. : Zhang (1984: 29, P. rusticola), Kalinina and Shabalin (2008). :,,,. :,,,,. : NK-PY,, WN, SK- (21 ) 150 180 1,000. (539ex): [SWU] : 46ex (,,,,, : v x.1981 1991). : 5ex (,,, : v viii.1984 1991). : 3ex (, : vii, viii.1979 1988). : 5ex (,,,, : v vii.1984 1994). : 5ex (13.vii.1985). [NAAS] : 1ex ( : 29.vii.1987). : 8ex ( : v ix.1922 1993). : 1ex ( : 26.v.1994); 2ex [Hakuyosan (= ): 25.vi. 1922]. : 1ex (vi.1919).?: 2ex (Kiho: 6.vi.1924); 1ex (Tashiro chikuto: 28.v.1982). [SNU] : 1ex ( : 30.ix.1965). : 32ex (,, : iv ix.1957 1994). : 2ex (, : v, vi.1988, 1995). : 1ex ( : 5.vi.1987). : 30ex (, : vi viii.1969 1988). [KU] : 17ex (,, : vi x.1971 79). : 1ex [Keijo (= ): 7.viii.1934]; 329ex (,,,,,,,, : v x.1955 1985). : 9ex (, : v viii.1957 1973). : 3ex (, : vii viii.1960 1974). : 15ex (,,,,, : v viii.1956 1991).?: 2ex (Zokuni: 15.viii.1930). [HSU] : 3ex ( : v vi.1994 1997). [HMNH-NK] : 14ex ( : 23.ix.1978). : 525, 4.5. 11.19. (iv- 1, v- 74, vi- 108, vii- 77, viii- 48, ix- 179, x- 28, xi- 10ex). :. 1990, 1997 1. : Okamoto and Nakayama, 1940: 198; Takahashi et al., 2006: 46 ( ); Kim, 1998: 351 ( ). Genus Glycyphana Burmeister, 1842: 345..,.,. : Cetonia horsfieldi Hope, 1831. in Grey, Zool. Miscell., p. 25. : 40 (Mikšić, 1982), 12 (Gorodinski, 2009) ( 1 ).

: : 57 :,. : Kim (2001b). 16. Glycyphana fulvistemma Motschulsky, 1860: 135 ( 2-16, 11-16, 17-16) Glycyphana fulvistemma Motschulsky, Schrenck s Reisen Amurl.: 135, 1860: Kolbe, 1886: 196; Heyden, 1887: 255; Bates, 1888: 377; Niijima and Kinoshita, 1923: 180; Okamoto, 1924: 180; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 91; Cho, 1957: 303; 1963: 220; 1969: 695; Cho et al., 1967: 198; 1968: 265; KSZ, 1968: 138 ( ); KSPP, 1972: 207; Kim and Kim, 1974b: 230; 1975: 255; Kim and Nam, 1982a: 157; 1982b: 277; Kim, 1978: 388; Stebnicka, 1980: 284; Kim, 1981: 345; 1993: 66; 2000a: 141; 2001: 132; 2001b: 150; 2002: 120; 2003: 127; 2004: 117; Mikšić, 1982: 390; ESK and KSAE, 1994: 154; Lie et al., 1997: 90. : (Cho, 1963), (Cho et al., 1967, 1968), (KSPP, 1972).,.,,.,.,., 3, 3. Ad: ( )..,,.. BL: 11 14 mm, BW: 6 8 mm. : Medvedev, 1952: 173; Zhang, 1984: 29. :,,,,. :,,. : NK-HB, HN,,,, PN, SK- (9 ) 100 130 400. (342ex): [SWU] : 3ex (,, : v vii.1984 1995). : 33ex (,,,, : iv ix.1975 1998). : 7ex ( : vi ix.2000 2003). : 2ex ( : vi, vii.1999); 1ex ( : 6.v.2002). : 7ex ( : 29.v.1999). : 1ex ( : 16.vii.2000). : 2ex ( : v vi.1991, 1992); 1ex ( : 22.v.1999). [NAAS] NK- : 3ex [Sharei (= ), Shakuoji (= ): vi vii.1922]. : 2ex [Kasan (= ): vi.1923]. 7ex ( : v vii.1958 1984). : 2ex (, : ix.1985, v.1987). : 1ex ( : 30.viii.1983). [SNU] : 3ex (, : v ix.1988 1997).

58 : 33ex (,,,,, : v x.1957 1995). : 3ex ( : vi vii.1983 1992). : 24ex (, : v vii.1969 1995). [KU] : 4ex (,,, : vi.1968 1977). : 133ex (,,,,,, : iv vii.1933 1992). : 2ex (, : v, vi.1977 1978). : 10ex (,, : v vii.1956 1991).?: 2ex (Kouryo: 12.ix.1933). [HSU] : 47ex ( : iv vi.1994 1997). [HMNH-NK] : 6ex ( : 2 5.vi.1991). : 1ex ( : 30.v.1970). : 286, 4.6. 10.10. (iv- 6, v- 128, vi- 122, vii- 17, viii- 4, ix- 8, x- 1ex). Genus Gametis Burmeister, 1842: 358.,,,.,.,,. 4 5 1. : Cetonia versicolor Fabricius, 1775. Syst. Ent., p. 51. : 6 (Mikšić, 1982), 6 (Gorodinski, 2009) ( 1 ). :,. : Kim (2001b). 17. Gametis jucunda (Faldermann), 1935: 386 ( 2-17, 11-17, 17-17) Cetonia jucunda Faldermann, Mem. Acad. Petrsb., II: 386, tab. 4, figs. 4, 5, 1835. Glycyphana jucunda: Kolbe, 1886: 195; Heyden, 1887: 255; Bates, 1888: 377. Oxycetonia jucunda: Niijima and Kinoshita, 1923: 182; 1937: 28; Okamoto, 1924: 180; Cho, 1934: 75; 1957: 303; 1963: 221; 1969: 696; Yoshino, 1935a: 14; Morita, 1936: 861; Masaki, 1936: 261; Doi, 1937: 62; 1938: 97; Mochizuki and Tsunekawa, 1937: 92; Nagaoka, 1938: 25; 1940: 474; Miwa and Chûjô, 1939: 92; Kim, 1956: 9; 1958: 98; 1960: 27; Ku, 1963: 28; Cho et al., 1967: 198; 1968: 265; KSZ, 1968: 138 ( ); Won and Choi, 1968: 379; Hyun and Woo, 1969: 193; Kim and Kim, 1971b: 160; 1972a: 84; 1974a: 109; 1974b: 230; 1975: 255; 1976: 102; Kim and Nam, 1977: 31; 1982a: 157; 1982b: 277; 1984a: 329; 1984b: 104; 1985: 105; Gu, 1973: 83; Kim, 1978: 389; Yoon and Nam, 1979a: 83; 1979b: 150; 1986: 160; Yoon et al., 1990: 111; Stebnicka, 1980: 286; Kim, 1981: 345; Nam and Kim, 1983: 129;

: : 59 Kim and Chang, 1983a: 167; 1984: 170; 1987b: 506; Park and Park, 1987: 673; Lee and Ko, 1988: 211; Lee et al., 1994: 147; Lie et al., 1997: 91. Gametis jucunda: Mikšić, 1982: 188; Kim et al., 1987a: 224; 1989: 177; 1991e: 180; 1992b: 154; 1994: 111; 1997a: 233; 1998: 170; 1999: 130; 2000a: 141; 2001: 132; 2002: 120; 2003: 127; 2004: 117; 2005: 80; 2006: 37; Yoon et al., 1990: 111; ESK and KSAE, 1994: 154; Kim, 1993: 66; 1995a: 166; 1995b: 140; 1996c: 174; 2001b: 151. Glycyphana kuperi Schaum, Trans. Ent. Soc. Lond., V: 69, 1848: Bates, 1888: 377. Oxycetonia jucunda kuperi: Niijima and Kinoshita, 1937: 28; Miwa and Chûjô, 1939: 92; Cho, 1957: 303. Oxycetonia jucunda ferruginosa Reitter, Best.-Tab., 38: 35, 1898: Niijima and Kinoshita, 1923: 182; Okamoto, 1924: 180; Kamijo, 1932: 21; Cho, 1934: 75; 1957: 303; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 92; Nagaoka, 1940: 474. Oxycetonia jucunda dolens Kraatz, Deut. Entom. Zeit., 23: 236, 1879: Cho, 1934: 75; Mochizuki and Tsunekawa, 1937: 92; Nagaoka, 1940: 474. Oxycetonia jucunda bealiae Gory-Percheron, Monogr. Ceton.: 282, 1833: Mochizuki and Tsunekawa, 1937: 92. Glycyphana albosetosa Motschulsky, Etud. Entom., X: 9, 1861: Okamoto, 1924: 180. Glycyphana argyroticta Burmeister, Hndb. Entom., 3: 360, 1842: Okamoto, 1924: 180. Oxycetonia Jucunda ab. borealis ( ): Nagaoka, 1940: 474. : (Cho, 1963 4), (Kim, 1958, 1960), (KSPP, 1972).,. 1. V,., 3. 1.. Ad:., 1.6.,,. BL: 10 14 mm, BW: 7 9 mm. : Murayama, 1931: 79; Medvedev, 1952: 174; Zhang, 1984: 28 (O. jucunda). :,,,,,,,,. :,,,. : NK-HB,,, HN,,,, PB (,, ), PN,, SK- (15 ) 140 200 7,000. (5,358ex): [SWU] : 167ex (,,,,,,,,, : iv viii.1980 2001). : 1,435ex (,,,,,,,,,,,,

60,, : iv x.1976 2000); 1ex ( : 25.vi.2005); 1ex ( : 26.viii.2005); 3ex ( : 15, 21.v.2002). : 66ex (,,,,,, : v ix.1986 2003); 1ex ( : 7.vi.2008). : 146ex (,,,,,,,,,, : v x.1979 2002); 1ex ( : 6.vii.2008). : 38ex (,,,, : v ix.1989 1993). : 6ex (v viii.1988 1994); 5ex ( : 27.v.2008); : 1,500ex. [NAAS] NK- : 20ex [Seishin (= ), Shuotsu (= ), Sharei (= ), Shakuoji (= ): v ix.1922, 1923]. : 1ex [Gesseiji (= ): 15.ix.1923]; 13ex (, : v x.1985 1995). : 4ex [Kasan (= ): v ix.1922]; 166ex (,,, : iv x.1922 1997); 4ex [Koryo (= ): iv vi.1923, 1924]. : 3ex (,, : v viii.1963 1991). : 106ex (,,,,, : v ix.1974 1993). : 14ex (,,,,,, : v ix.1950 1996). : 25ex (iv x. 1919 1996). [SNU] : 1ex ( : 11.v.1985). : 1ex ( : 20.vi.1988); 486ex (,,,,,, : iii x.1969 1997). : 7ex (, : v vi.1984 1992). : 4ex ( : vi.1987). : 28ex (, : v viii.1969 1997). : 5ex iv ix.1967 1992). [KU] : 18ex (,,,, : iv viii.1978 1988). : 412ex (,,,,,, : iv x.1969 1992). : 29ex (,, : vi viii.1978 1990). : 7ex ( : vii.1978). : 20ex (,,,, : v vii.1977 1991). : 24ex (4.v.1978). [HSU] : 245ex ( : iv vi.1994 1997). [HMNH-NK] : 17ex ( : 2 5.vi.1991). : 2ex ( : 16.vii.1982). : 6ex (Kyollyong Reservoir: 30.ix.1978); 2ex ( : 1.x.1978); 11ex ( : 1.x.1978); 1ex ( : 23.v.1970); 6ex (16.viii.1971); 1ex ( : 5.ix.1971). : 1ex ( : 18.viii. 1971); 61ex (2.ix.1971); 1ex ( : 29.ix.1979); 4ex (31.viii.1971); 1ex ( : 11.ix.1980); 133ex ( : 10.vi.1970). : 22ex ( : 29, 30.v.1970); 1ex (12.vii.1977); 1ex (17.vi.1985); 3ex (12.vii.1991); 2ex ( : 29.ix.1979); 1ex (Chonjon: 9.vi.1991). : 9ex ( : 28.ix.1978); 7ex ( : 27,28.ix.1978). : 17ex ( : 6 8.vi.1970). : 3,430 3.20. 11.3. (iii- 2, iv- 148, v- 902, vi- 1,156, vii- 221, viii- 174, ix- 638, x- 84, xi- 1ex). : Nakayama and Okamoto, 1940: 199; Saito, 1941: 12. :.. Mikšić (1982) 13 5. ( ) Tribe Cremastocheilini Burmeister and Schaum, 1841: 243. Cremastochilidae Burmeister and Schaum, 1841: 243...

: : 61.. : Cremastocheilus Knoch, 1801: 115. : 2 (Gorodinski, 2009), 4 (Arnett, 1963) ( 1 ). : 55 (Gorodinski, 2009), 36 (Arnett, 1963) ( 1 ). :,,,. : Krikken (1977), Ross (1963). Genus Clinterocera Motschulsky, 1858: 112. Callynomes Westwood, 1874. Cholerastoma Mohnike, 1871. 2 Stebnicka (1980) 1. : Cremastochila scabrosa Motschulsky, 1853: 47. Clinterocera Medvedev, 1964: 334 341 (6 species). 18. Clinterocera obsoleta (Fairmaire), 1878: 108 ( 2-18, 11-18, 17-18) Callynomes obsoletus Fairmaire, Ann. Soc. Ent. Fr., (5) VIII: 108, 1878: Kolbe, 1886: 196; Niijima and Kinoshita, 1923: 185; Murayama, 1935b: 6; Mochiuzuki and Tsunekawa, 1937: 96; Miwa and Chûjô, 1939: 94; Cho, 1957: 303; 1969: 696; KSZ, 1968: 138 ( ); Kim and Kim, 1974b: 230; 1975: 256; Kim and Nam, 1982a: 157; 1982b: 277; Kim, 1981: 345; Kim et al., 1987b: 506; 1998c; 357; 2004: 117. Clinterocera obsoleta: Stebnick, 1980: 210; Yoon et al., 1990: 111; ESK and KSAE, 1994: 154 ( ). Clinterocera mandarina Westwood, 1874 ( ): Stebnicka, 1980: 210 (, HB;, HB;, WN); Kalinina, 1989: 432; Lie et al., 1997: 92. : (ESK and KSAE, 1994). :., 1...

62, (mentum).,... 2. 7. Ad:,,,. 90. BL: 15 17 mm, BW: 4 5.5 mm. :,. :,,. : NK-HB, PB, PY, SK-,,,,,, GS,,,,,,,,,,,,,,,,,,,,,,,,,,,, (,, ),, CBN,,,,,, GBN,,,,,,,,,,, JBN,,. (201ex): [SWU] : 6ex (, : v vii.1981 1987). : 82ex (,,,,,,,, : iv ix.1975 1996). : 6ex (,,, : v, vi.1989, 1995, 1999, 2003). : 13ex ( : v.1996); 1ex ( : 28.v.1999). : 1ex ( : 4.vii. 1993). [NAAS] : 1ex ( : 30.v.1982). : 1ex [Koryo (= ): 1.vi.1924]; 1ex ( : 13.v.1990). : 1ex ( : 18.vii.1993). [KU] : 14ex (,, : v vi.1967 1984). : 51ex (,,,,,,,, : iv viii.1932 1992). : 2ex (, : v.1984, 1987); 1ex ( : 23.vii.1931). : 11ex (, : iv vi.1974 1989). : 3ex (, : v vii.1985 1991).?: 1ex (Gunnaido Cosho-c: 28.iv.1937). [HSU] : 1ex ( : 17.v.1997). : 201, 4.5. 9.20. (iv- 6, v- 118, vi- 67, vii- 3, viii- 4, ix- 3ex). Family Rutelidae MacLeay, 1819: 69. Rutelinae MacLeay, 1819.,.,,,,.,. 9 10, 3.,. (, terminal spur) 2. 2. (, Onychium).,. (Phytophagous),., 2,

: : 63. (Saprophagous). (Rutelidae) (Melolonthidae) Machatschke (1972) Lawrence and Newton (1995) (Scarabaeidae).. 6 ( ), (Rutelini) (Anomalini) (Neotropic Region), ( ), (Adoretini). Machatschke (1972) (Rutelidae) 200 4,100 5,000,. : Rutela Latreille, 1802: 151. : Reitter (1903), Ohaus (1915a, 1915b), Murayama (1938a), Medvedev (1949), Machatschke (1972), Kalinina (1989), Kim (1995c, 1995d, 1996a, 1996b, 1997a, 1998a, 1998b, 2001b), Liu et al. (1997), Zorn (2004); : Murayama (1931), Medvedev (1952), Zhang (1984). (Rutelidae) : Kolbe (1886) 6 1. ( ). Heyden (1887) 10, 1990, 16 80, 20. 1990 (Kim, 1995c, d, 1996a, b, 1997a, 1998a, b) 69 (Adoretini), (Rutelini), (Anomalini) 11 35. 4. : 6 ( 3 ). : 200 250 (?), 200 (Machatschke, 1972) ( 11 ). : 4,500 5,000 (?), 4,100 (Machatschke, 1972) ( 35 ). :. : Kolbe (1886), Heyden (1887), Reitter (1903), Ohaus (1915a, 1915b), Nijiima and Kinoshita (1923, 1927), Murayama (1938a), Medvedev (1949), Kalinina (1989), Kim (1995c, 1995d, 1996a, 1996b, 1997a, 1998a, 1998b, 2001b). 1.., 10. 3, 2 tribe Adoretini 2 2. 10. 3, (, inner spur)

64.. (pleural suture) Tribe Rutelini 9. 2,.. Tribe Anomalini Tribe Adoretini Burmeister, 1844: 466. Adoretidae Burmeister, 1844: 466. Adorrhinyptiina, Adoretina, Pachyrhinadoretina, Trigonostomina, Prodoretina 5, Adoretina Adoretus, Adorodocia, Adoroleptus, Pseudadoretus, Scaphorhinadoretus 5, Adoretus 100. : Adoretus Dejean, 1833. : 5 ( 1 ). : ( 2 ). :. : Kobayashi (1937), Kuijten (1922), Sawada (1938), Kim (1995c). Genus Adoretus Dejean, 1833: 157. 5 1, 2 2. : Melolontha nigrifrons Steven, 1809. by subseq. design. Medvedev, 1949: 313 (from Russia and Central Asia). : 100 ( 2 ). :,,. 1.,

: : 65 Lepadortus,.., tenuimaculatus, Chaetadoretus.. hirsutus 19. Adoretus tenuimaculatus Waterhouse, 1875: 112 ( 2-19, 11-19, 17-19) Adoretus tenuimaculatus Waterhouse, Trans. Ent. Soc. Lond., 1875: 112 ( Lepadoretus ): Kolbe, 1886: 192; Bates, 1888: 376; Niijima and Kinoshita, 1923: 141; Okamoto, 1924: 178; Maruta, 1929: 370; Yoshino, 1935a: 14; Morita, 1936: 861; Masaki, 1936: 260; Mochizuki and Tsunekawa, 1937: 90; Cho, 1957: 295; 1963: 219; 1969: 682; Cho et al., 1967: 197; 1968: 265; Hyun and Woo, 1969: 197; Kim and Kim, 1971: 160; 1972a: 84; 1974a: 230; 1975: 250; Kim and Nam, 1982a: 156; 1985: 105; Kim, 1978: 369; Yoon and Nam, 1979: 149 ( ); 1990a: 140; 1990b: 110; Kim, 1981: 345; 1992: 106; 1993: 65; 1995a; 165; 1995b: 139; 1995c: 38; 1997: 231; 2001: 83; Kim et al., 1982a: 145; 1987c: 506; 1991a: 232; 1991b: 68; 1991c: 154; 1991d: 192; 1991e: 179; 1992b: 154; 1994: 111; 1996a: 58; 1996b: 128; 1998: 170; 1999: 130; 1996b: 128; 1998: 170; 1999: 130; 2000: 132; 2001: 132; 2001b: 83; 2002: 120; 2003: 126; 2004: 117; 2005: 79; 2006: 36; Nam and Kim, 1983: 128; Park and Cho, 1986: 128; 1992: 177; Park and Park, 1987: 673; Park and Kim, 1993: 113; Lee et al., 1994: 147; ESK and KSAE, 1994: 152; Lie et al., 1997: 46. Adoretus umbrosus var. tenuimaculatus: Heyden, 1887: 254; Okamoto, 1924: 178. Adoretus falciungulatus Nomura, 1965 ( ): Kim and Chang, 1987c: 506. Adoretus sinicus Burmeister, 1885 ( ): Nomura, 1965: 8; KSPP, 1972: 205; Stebnicka, 1980: 280 ( ). : ( ) (Cho, 1963 13); (Cho et al., 1967); (Hyun and Woo, 1969); (Ku, 1973; Kim et al., 1985); (Kim, 1978 5). : ( )..,.,.,,.,. 10. Ad: ( ),

66. BL: 8.5 14 mm, BW: 4.5 7.5 mm. :,,,. :,,,. : NK-HN, PY, SK- (12 ) 130 180 4,000. (3,280ex): [SWU] : 64ex (,,,,,,,,,,, : v viii.1981 2002); 1ex ( : 10.vii.2009). : 500ex [,,, (4 ) : v x.1961 2003]; 3ex ( : 12.vi.2005). : 173ex (,,,,,,,,,,,,,,,, : v x.1986 2003). : 75ex (,,,,,,,,,, : v i.1978 1998); 2ex ( : 6, 7.vi.2008). : 1ex ( : 6.vii.2008). : 117ex (,,,,,,,, : v x.1979 2002). : 1ex (viii.1988); 2ex (5.v.1997). [NAAS] : 11ex (,,, : v ix.1974 1997). : 152ex (,,,,,,,, : iv x. 1921 1998). : 60ex (,,,,, : v viii.1983 1997). : 30ex (,,,,,,,,, : v viii.1976 1997). : 22ex (,,,,,, : vi xi.1921 1996). : 14ex (iv x.1975 1996). [SNU] : 9ex (,, : v viii.1977 1995). : 314ex [,,, (5 ),,,,,,,, : iv x.1957 1997]. : 41ex (,,,,,, : iii xi.1984 1995). : 18ex (,,,,, : vi viii.1987 1996). : 189ex (,,,, : v ix.1968 1997). : 3ex (8.viii.1997). [KU] : 30ex (,,,, : iv ix.1968 1992). : 524ex [,,,,,, (5 ),,,, : iv x.1929 1996]. : 8ex (,,,, : v viii.1973 1987). : 84ex (,,,,, : vi vii.1974 1996). : 151ex (,,,,, : iv ix.1972 1993). : 21ex (v vii.1966 1990). [EWU] : 5ex (, : vi.1973, 1979). ; 199ex (,,,,,,,,, : iv x. 1956 1993). : 7ex (,, : vii viii.1971 73). : 15ex (, : vi vii.1978 1986). : 1ex ( : vii.1975). : 35ex (v vi.1965 1973). [HSU] : 34ex ( : iv vi.1994 1997), [HMNH-NK] 2ex (, : 4 6.viii.1975); 1ex (9 11.vii.1977). : 6ex (Kyollong Resevoire: 30.ix.1978); 1ex ( : 16.vi.1988); 3ex ( : 15.vi.1988). : 4ex ( : 13 18.vii.1982); 2ex (12 28.v.1991); 1ex ( : 1.x.1978); 2ex ( : 13.ix.1979); 1ex ( : 25.viii.1989). : 1,667, 4.21. 11.28. (iv- 10, v- 402, vi- 787, vii- 222, viii- 103, ix- 110, x- 18, xi- 14ex). : Saito, 1931, 1941; Kamijo, 1936; Nakayama and Okamoto, 1940; Murayama, 1938; Kim and Kang, 1993: 415; Lee et al., 1997: 156 165; 2002: 217 223.

: : 67 20. Adoretus hirsutus Ohaus, 1914: 503 ( 2-20, 11-20, 17-20) Adoretus hirsutus Ohaus, Deut. Ent. Zeits., 1914: 503; Murayama, 1935: 5; 1937a: 35; Cho, 1957; KZS, 1968: 136; 295; Kim and Lee, 1991b: 68; Kim, 1992: 106; 1993: 65; 1995c: 39; 2001b: 85; ESK and KSAE, 1994: 152 ( ). : (KZS, 1968). :..,..,.. 2.5,.,. 2. Ad: ( ). BL: 9 11.5 mm, BW: 4.5 6 mm. : Zhang, 1984: 43. :,,,. :,. : SK-GS,,,,,,,,,,,,,,,,,,,,, CB,.,, CN,,,, GB,,,.,,, GN,, JB, JN,,. (214ex): [SWU] : 1ex ( : 10.vii.2009). : 41ex (,,,, (14 ),,,,, : vi ix.1972 1998); 1ex ( : 4.vii.2003). : 8ex (,,,, : v vii.1979 1993). : 10ex (,,,, : vi viii.1986 1996); 1ex ( : 19.vii.2001). : 1ex ( : 26.vii.1996). : 2ex (, : vii.1996, 1998); 1ex ( : viii.1996). [NAAS] : 94ex ( : vi vii.1958 1996). : 4ex ( : vii.1975). [SNU] : 1ex ( : 25.v.1996); 1ex ( : viii.1995); 94ex ( : vi vii.1958 1996). : 4ex ( : vii.1975). [SNU] : 1ex ( : 25.v.1996); 1ex ( : 4.viii.1995); 2ex ( : viii ix.1995). [KU] : 7ex (,, : vi, vii.1983 1994). : 6ex (,,, : vii ix.1967 1984). : 1ex ( : 8.vii.1976). [EWU] : 2ex ( : viii, ix.1973, 1974). : 26ex (,,,,, : vi x.1960 1984). : 1ex ( : 3.viii.1964). : 1ex ( : 21.vii.1966). [HMNH-NK] : 1ex ( : 8.vii.1982). : 1ex ( : 16.vii.1982). : 212, 4.2. 9.17. (vi- 15, vii- 149, viii- 45, ix- 3ex). : Murayama, 1938a: 259 264; Kim and Kang, 1993: 415.

68 Tribe Rutelini MacLeay, 1819: 69. Rutelidae MacLeay, 1819.. 10.. 3, (, inner spur). 7 (pleural suture).,,. : Rutela Latreille, 1802: 151. : 70, Parastasia 1 ( 1 ). : ( 1 ). :. : Reitter (1903), Ohaus (1915a), Murayama (1938a), Medvedev (1949), Kalinina (1989), Kim (1995c). Genus Parastasia Westwood, 1842: 204. : Parastasia canaliculata Westwood, 1841 by subseq. design. Machatschke, 1972. : ( 1 ). :,,,. 21. Parastasia ferrieri Nonfried, 1895: 289 ( 3-21, 11-21, 18-21) Parastasia ferrieri Nonfried, Berl. ent. Zeit., 40(3): 289, 1895: Kim, 1995c: 40; 2001b: 86. Ohkobous quadridentatus Sawada, Nippon no Kôchû, 2(2): 74, 1938. Ohkubous ferrieri quadridentatus: Kim and Kim, 1972b: 197; ESK and KSAE, 1994: 152 ( ). : (Loo-Choo, Amami-Ôsima)..,.

: : 69. 4, 2. 2,.. 3,. Ad:,. BL: 11 14 mm, BW: 7 8 mm. :,, (Ryukyu). :,. : SK-GG,,, GB,, GN (, ),, JN. (13ex): [SWU] : 3ex ( : 10.viii.1999, UV light, Kim); 1 ( : 8.vii.1981, E.Y. Lee); 1 ( : 17.viii.1993, Kim). : 1ex ( : 13.viii.1999); 1ex ( : 10. viii.1999, UV light, Kim). : 2ex ( : 29, 30.viii.1981, UV light, Kim); 1 (8.viii.1989; Jeon); 1ex ( : 8.viii.1985); 1 ( : 5.viii.1960, Cho). : 1 ( : 15.viii.1972, Kim). Tribe Anomalini Streubel, 1839: 136. Anomalidae Streubel, 1839., 9. ( ). 3,. 7.,. (Popillina) (Anomalina), Popillia 23 4. 1,000, Anomala 11 72 8 28. : Anomala Samouelle, 1819: 191. : ( 9 ). : ( 28 ). :. : Kolbe (1886), Heyden (1887), Reitter (1903), Ohaus (1915a), Murayama (1938a), Medvedev (1949), Machatschke (1972), Kalinina (1989), Kim (1995d, 1996a, 1996b, 1997a, 1998a, 1998b, 2001); internet: Popillia-Wikipedia, the free encyclopedia, 2009; : Ritcher (1966).

70 1., (subtr. Popillina) Popillia, subtr. Anomalina Subtribe Popillina Ohaus, 1918: 133. Popillina Ohaus, 1918. : Popillia Dejean, 1821: 60. Genus Popillia Dejean, 1821: 60. Calopopillia Kolbe, 1894 ( : candezei Kraatz, 1892). Eupopillia Kolbe, 1894 ( : callipyga Dohrn, 1879). Metapopillia Kolbe, 1894 ( : biimpressa Kraatz, 1892). Peripopillia Kolbe, 1894 ( : oberthüri Kraatz, 1892). Pseudopopillia Kolbe, 1894 ( : phylloperthina Kraatz, 1892). Xenopopillia Kolbe, 1910 ( : ducatrix Kraatz, 1892). Godschama Reitter, 1903 ( : hexaspila Ancey, 1883). Costapopillia Benderitter, 1922 ( : callewaerti Benderitter, 1922). Popilia. (P. japonica=japanese beetle). 1916 New Jersey Riverton Canada. 23, 4 1. : Trichius bipunctatus Fabricius, 1787. Mant. Ins., II, Append. : 130, 90 (Loebl and Smetana, 2006) ( 4 ).

: : 71 :,, 1 (P. japonica). : Kolbe (1886), Heyden (1887), Kraatz (1892), Medvedev (1949), Lin (1988), Kim (1995d, 2001b). 1. 10 15 mm,. 2 6 12 mm,. 3 2.. 1 flavosellata,. mutans 3... 1 quadriguttata a.,.,. q. quadriguttata b.,.,. q. japonica,.. 1 formosana 22. Popillia flavosellata Fairmaire, 1886: 331 ( 3-22, 11-22, 18-22) Popilia atrocoerulea Bates, Proc. Zool. Soc. Lond., 1888: 376 (Popillia, atrocaerulea, atsocoerulea ): Kraatz, 1892: 245; Reitter, 1903: 46; Niijima and Kinoshita, 1923: 133; Okamoto, 1924: 178; Kamijo, 1932: 21; Cho, 1934: 75; 1957; 1963: 294 ( ); 1969: 682; Cho et al., 1968: 265; Masaki, 1936: 261; Morita, 1936: 861; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 48; Kim and Kim, 1974a: 230; 1974b: 108; 1975: 248; 1976: 102; 1977: 130; Kim and Nam, 1982a: 156; 1982b: 276; 1983: 328; 1984: 87; 1985: 105; Kim, 1978: 366; Yoon and Nam, 1978: 83; 1990: 110; Kim et al., 1979: 84; 1983: 166; 1987a: 104; 1987b: 224; 1987c: 506; 1994: 111; Kim, 1981: 345; 1995d: 212; Stebnicka,

72 1980: 271; Shin and Park, 1981: 131; Nam and Kim, 1983: 129; Park and Cho. 986: 128; 1992: 178; Park and Park, 1987: 673; Lee et al., 1994: 147. Popillia atrocoerulea atra Kraatz, Deut. Ent. Zeit., 2: 245, 1892: Reitter, 1903: 46. Popilia coerulea Boheman, Resa Eugen.: 55, 1858: Kolbe, 1886: 190; Kraatz, 1892: 245. Popilia adamas Heyden, Horae. Soc. Ent. Ross., 52: 190, 1887 (nec. Newman): Kraatz, 1892: 245. Popilia flavosellata Fairmaire, Ann. Soc. ent. Fr., 6(6): 331, 1886. Popillia flavosellata: Kim et al., 1991b: 68; 1991d: 192; 1991e: 179; 1992: 154; 1996b: 128; 1999: 130; 2000: 132; 2002: 120; 2004: 117; 2005: 79; Kim, 1992: 106; 1993: 65; 1994: 214; 1995a: 165; 1995b: 140; 1996c: 174; 1997: 231; 1998: 170; 1999: 130; 2001b: 89; Lin, 1988: 27; ESK and KSAE, 1994: 153; Lie et al., 1997: 34. : (Cho, 1969 14); (KSPP, 1972; Ku, 1973).,... 1/3,.,.. Ad:,, (P. mutans).,. BL: 10 15 mm, BW: 6 9 mm. : Murayama, 1931: 73. :,, (Primorskij),. :,,,. : NK-HB,,, HN, PB, PY,, GW, SK- (15 ),, 130 150 5,000. (2,450ex): [SWU] : 269ex (,,,,,,,,,,,,,, : iv viii.1981 2000). : 225ex [,,,,,,,,, (4 ),,,,, : iv x.1961 2003]. : 71ex (,,,,,,, : vi viii.1986 2003). : 78ex (,,,,,,,,,,,, : iv ix.1979 2000). : 22ex ( : vi.1990, 1991); 43ex (,,,,,,, : v x.1982 1998). [NAAS] : 44ex (,,,, : vi viii.1987 1997). : 8ex (,,, : vi viii.1973 1997). : 1ex ( : 16.vii.1996); 2ex ( : 25.vi.1997). : 3ex ( : 26.vi.1973). [SNU] : 1ex ( : 20.vii.1997). : 66ex (,,,,,,,,, : iv ix.1922 1997). : 13ex (,, : vi xi.1983 1992). : 58ex (,,,,, : vi viii.1968 1997). : 2ex (vii, viii.1971, 1997). [KU] : 109ex (,,,, : v vii.1967 1994). : 416ex [,,,,,,, (3 ),,, : iv ix.1931 1991]. : 59ex (,,,,, : v viii.1957 1992).

: : 73 : 305ex (,,,,, : v vii.1974 1990). : 130ex (,,,, : iv x.1970 1993). : 2ex ( : 2.viii.1974). [EWU] : 104ex (,,,,, : v viii.1962 1984). : 314ex (,,,,,,,,, : v x.1957 1998). : 14ex (,, : vii, viii.1969). : 17ex (,,,,,, : vii viii.1959 1986). : 44ex (,,,,,,, : v viii.1969 1992). : 3ex (v viii.1971 1982). [HSU] : 18ex ( : v vii.1994 1997). [HMNH-NK] : 4ex ( : 2 7.vi.1991). : 5ex ( : 12.vii.1991). : 1,544, 4.24. 10.29. (iv- 3, v- 47, vi- 849, vii- 601, viii- 37, ix- 4, x- 3ex). : 1998 6 19 ( ).,. 23. Popillia mutans Newman, 1838: 337 ( 3-23, 11-23, 18-23) Popillia indigonacea Motschulsky, Etud. Entom., II: 47, 1853 (Popilia ): Kolbe, 1886: 190; Bates, 1888: 376; Niijima and Kinoshita, 1923: 134; Okamoto, 1924: 178; Kamijo, 1932: 21; 1936: 85; Morita, 1936: 861; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 49; Doi, 1937: 63; 1938: 3; Nagaoka, 1940: 474; Kim, 1956: 9; 1958: 98; 1960: 27; Cho, 1957: 294; 1963: 220 ( ); 1969: 681; Hyun and Woo, 1970: 80; Kim and Kim, 1972b.: 197; 1974a: 230; 1974b: 108; 1975: 249; 1976: 102; 1977: 130; Kim and Nam, 1978: 134; 1982a: 156; 1982b: 276; 1984b: 328; Kim, 1978: 366; Shin, 1979: 144; Kim et al., 1979: 84; 1982a: 145; 1982b: 173; 1983: 166; 1984: 169; 1987a: 104; 1987b: 224; 1987c: 506; Kim, 1981: 345; Yoon and Nam, 1979: 149; 1990a: 140; 1990b: 110; 1985: 160; Stebnicka, 1980: 270; Lee and Kwon, 1981: 154; Park and Cho, 1986: 128; 1992: 178; Lee et al., 1994: 147; ESK and KSAE, 1994: 153. Popilia mutans Newman, Ann. Mag. Nat. Hist., 2(II): 337, 1838 (Popilia ): Kim et al., 1991b: 68; 1991d: 192; 1991e: 179; 1992: 154; 1996a: 58; 1996b: 128; 1998: 170; 2000: 132; 2001: 132; 2002: 120; 2003: 126; 2004: 117; 2006: 37; Kim, 1992: 106; 1993: 65; 1994: 214; 1995a: 165; 1995b: 1995d: 210; 1996c: 174; 140; 1997: 231; 2001b: 87; Lin, 1988: 15; ESK and KSAE, 1994: 153; Lie et al., 1997: 32. Popillia mutans indigonacea: Kraatz, 1892: 240. : (KSPP, 1972 9). : (Peiking).,. (flavosellata),.,.

74.,.. Ad:,.. BL: 10 15 mm, BW: 6 9 mm. : Zhang, 1984: 47. :,,,,. :,,,,. : NK-HB, HN,,, PB,, PN,,,, WH, GW, SK- (10 ) 120 150 3,000. (2,079ex): [SWU] : 30ex (,,,,,,,,,, : vii xii.1981 2001). : 875ex (,,,,,,,, : iv xi.1944 2002). : 85ex (,,,,,,,,,,,,, : vi ix.1977 2002). : 63ex (,,,,,,,,,,,,,,,,, : vi ix.1979 1998); 2ex ( : 19, 20.vii. 2001). : 143ex (,,,,,,,,,,,,,,, : vi x.1980 1998). : 12ex (viii x.1983 1988). [NAAS] : 25ex ( : viii.1987). : 11ex ( : vi ix.1976 1995). : 1ex ( : 25.vii.1990); 2ex ( : viii.1996). : 1ex ( : 13.vii.1998); 6ex ( : viii.1993). : 1ex ( : 20.viii.1992). : 52ex (vii ix.1987 1995). [SNU] : 4ex (,, : vi ix.1961 1992). : 134ex [,,, (4 ),,,,,, : iii x.1916 1997]. : 16ex (,,,,, : iii ix.1960 1996). : 16ex (,, : viii.1988 1996). : 7ex (, : viii.1995, 1996); 23ex (, : vi viii.1959 1997). : 3ex (viii.1994 1997).?: 6ex ( : viii.1993). [KU] NK-2ex ( : vii, viii.1932, 1933); 2ex ( : vii.1933, 1938). : 8ex (,,,, : vi viii.1967 1984). : 115ex (,,,,,,,, : iv x.1921 1991). : 111ex (,,,,, : vii viii.1954 1993). : 35ex (,,,,,, : vi viii.1974 1990). : 26ex (,,,,,,, : iv viii.1955 1995). : 15ex (vii viii.1959 1990).?: 3ex ( : 28.vii. 1993). [EWU] : 15ex (,,, : vii viii.1958 1980). : 171ex (,,,,,,,,,, : v x.1956 1984). : 2ex (, : vi, vii.1967, 1969). : 7ex (,,, : vii viii.1956 1983). : 20ex (,,,,,,, : vi viii.1961 1992). [HSU] : 10ex ( : v vii.1994 1997). [HMNH-NK] : 2ex ( : 13, 26.ix.1978); 2ex ( : 3.vii.1988). : 2ex ( : 31.vii. 1882). : 6ex [ (3 ): vii, viii.1975, 1991]. : 1,508, 4.3. 12.15. (iv- 5, v- 4, vi- 27, vii- 282, viii- 927, ix- 211, x- 41, xi- 8, xii- 3ex).

: : 75 24. Popillia quadriguttata (Fabricius), 1787: 377 ( 3-24, 11-24, 18-24) Trichius quadriguttata Fabricius, Mant. Ins., II, Append.: 377, 1787. Popillia quadriguttata: Stebnicka, 1980: 272; Kim, 1992: 106; 1993: 65; 1995a: 165; 1995b: 140; 1995d: 215; 2001: 91; Lin, 1988: 24; ESK and KSAE, 1994: 153; Ku et al., 1999: 171; Lee et al., 2002: 17. Popillia quadriguttata quadriguttata: Kalinina, 1989: 411; Kim, 1995d: 214 ( ); 2001b: 91. Popillia quadriguttata japonica Kim, Kor. J. Entom., 25(3): 215, 1995d (, syn. nov.): 2001b: 91. Popillia japonica Newmann, Ann. Mag. Nat. Hist., (2)II: 337, 1838 (Popilia )(?): Heyden, 1887: 253; 1889: 662; Yoshino, 1935a: 14; Kim, 1956: 9; 1958: 98; 1960: 27; Ku, 1963: 28; Cho et al., 1967: 198; Cho, 1969: 682; Hyun and Woo, 1969: 194; Kim and Kim, 1975: 249; Kim and Nam, 1982a: 156; 1982b: 276; 1984b: 328; 1985: 105; Kim, 1978: 368; Yoon and Nam, 1978: 83; 1979: 149; 1985: 160; 1990b: 110; Kim et al., 1979: 84c; 1987b: 224; 1987c: 506; 1991a: 68; 1991b: 192; 1991e: 179; Stebnicka, 1980: 272; Park and Cho, 1986: 128; 1992: 178; Park and Park, 1987: 673; Lee and Ko, 1988: 211; Lee et al., 1994: 147; ESK and KSAE, 1994: 153; Kim, 1992: 154; 1995d: 215; Lie et al., 1997: 32; Ku et al., 1999: 171. Popilia bogdanowi Ballion, Bull. Soc. Hist. Nat. Moscou, IV: 345, 1870: Bates, 1888: 376; Miwa and Chûjô, 1939: 48; Cho, 1957: 294. Popillia chinensis Frivaldszky, Term. Füz., XII(4): 201, 1889: Kraatz, 1892: 249; Reitter, 1903: 47; Arrow, 1913: 39; Stebnicka, 1980: 208. Popillia chinensis sordida Kraatz, Deut. Ent. Zeit., II: 249, 1892. Popillia uchidai Niijima and Kinoshita, Bull. Exp. For., Hokkaido Univ., 2: 133, 1923: Kamijo, 1932: 21; Morita, 1936: 861; Doi, 1937: 63; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 50; Nagaoka, 1940: 474; Cho, 1957: 295; 1963: 220; Cho et al., 1968: 265; Kim and Kim, 1974a: 230; Stebnicka, 1980: 208; Kim and Chang, 1987c: 506; Kim and Lee, 1991b: 68; ESK and KSAE, 1994: 153; 2005: 79; Kim, 1995d: 215 (syn. nov.). Popillia ruficollis Kraatz, Deut. Ent. Zeit., II: 254, 1892: Stebnicka, 1980: 208 ( ). : japonica= (Kim, 1958 17), ( ) (Kim, 1960; Cho et al., 1967); uchidai= ( ) (Cho, 1963 4), (KSPP, 1972); bogdanowi= (KZS, 1968); quadriguttata= (ESK and KSAE, 1974).,.,...,., 1.. Ad: (flavosellata),.

76,,,. q. quadriguttata=bl: 8 11 mm, BW: 4.5 6.2 mm; q. japonica=bl: 10 12 mm, BW: 6 7 mm. : Murayama, 1931: 71 (P. uchidai); Zhang, 1984: 47. : Lee et al., 2002: 15 19; 2007: 103 109. :,,,. : q. japonica=, ; q. quadriguttata=,,. : q. japonica=nk-hb, SK-GG ; q. quadriguttata=sk- (6 ) 100 130 1,000. : q. japonica (2ex): [SWU] : 1 ( : 1921, P.S. Cho). : 1 ( : 10.vi.1934, Eguti). q. quadriguttata (874ex): [SWU] : 39ex (,,,,,, : vi viii. 1981 1998); 1ex ( : 9.vii.2006). : 143ex [,,,,,,,,, (25 ),,,, : vi x.1934 1998]. : 42ex (,,,,,,,,, : v viii.1974 1997). : 66ex (,,,,,,,,, : vi x.1974 2000). : 9ex (,,,, : vi viii.1984 1995). : 7ex (viii.1972 1991). [NAAS] : 1ex ( : 5.viii.1987). : 31ex ( : iv viii. 1974 1997); 1ex ( : vii.1993). : 16ex ( : vii.1987). : 1ex ( : 16.vii.1996). : 2ex ( : 17.vii.1996); 1ex ( : 21.ix.1992). : 54ex (vi viii.1983 1995). [SNU] : 4ex ( : viii ix.1993 1995). : 102ex (,,,,,,, : iv x.1955 1997). : 31ex (, : vi ix.1983 1996). : 1ex ( : 24.vi.1989). : 5ex (, : vi vii. 1969 1988). : 13ex (vii viii.1965 1995). [KU] : 8ex ( : vi vii.1931 1984). : 29ex [,,,, (3 ),,, : vi vii.1931 1989]. : 6ex (,, : vii, viii.1979, 1982). : 57ex (,,,, : vii.1974 1990). : 16ex (,, : vi vii.1960 1985). : 11ex (vii viii.1932 1990). [EWU] : 8ex (,, : vi viii.1967 1975). : 92ex (,,,,,,, : iv x.1954 1983). : 10ex (,,, : vii, viii.1969 1971). : 7ex (, : vi vii.1969 1975). : 11ex (,,, : vi ix.1965 1975). : 24ex (vii viii.1957 1979). [HMNH-NK] : 2ex ( : 17.viii.1982). :( : 20.vii.1982); 1ex ( : 3.vii.1988); 1ex ( : 1.vii.1991). : 20ex (Tesson waterbasin: 4.vii.1977). : 661, 4.19. 10.16. (iv- 3, v- 15, vi- 116, vii- 410, viii- 89, ix- 19, x- 7ex). 25. Popillia formosana Arrow, 1913: 45 ( 3-25, 11-25, 18-25) Popillia formosana Arrow, Ann. Mag. Nat. Hist., 8(12): 45, 1913: Kim, 1995d: 215. Popillia quelpartiana Ohaus, Stett. Entom. Zeit., 84: 172, 1924: Stebnicka, 1980: 208; ESK and KSAE,

: : 77 1994: 153; Kim, 1995d: 215 (syn. nov.); 2001b: 92. : (Tai-nan)....,.,.. Ad:,,., 2/3.. BL: 6 9 mm, BW: 4 5 mm. :,,. :,. : JN: (, ), JJ:,,,. (19ex): [SWU] : 1 1 [ (, ): 6.vii.1982, Kim]. : 1 ( : 15.vii.1985); 1 2 ( : 2.viii.1955); 1 ( : 4.viii.1972, Kim); 11ex ( : 12.vii. 1983, Kim); 1 (18.vii.1966). Subtribe Anomalina Streubel, 1839: 136. Anomalidae Streubel, 1839., 9,,. Anomala 1,000. 11 72 (Kim, 1995d 2001b) 8 28. : Anomala Samouelle, 1819: 191. : ( 8 ). : ( 28 ). :. : Heyden (1887), Reitter (1903), Ohaus (1915a, 1915b, 1943), Murayama (1938a), Sawada (1938, 1941), Medvedev (1949), Machatschke (1952), Kalinina (1989), Kim (1995d, 1996a, 1996b, 1997a, 1998a, 1998b, 2001b).

78 1.. 2, 1. 3 2.,.,, Proagopertha,,. 1 Phyllopertha 3.. Mimela, 4 4., Callistethus 5 5. 2 4.. 13 mm Blitopertha 2 4. 12 mm 6 6.,. Chejuanomala 7 7. 10 Bifurcanomala Anomala Genus Proagopertha Reitter, 1903: 50. Reitter (1903) Anomala pubicollis Waterhouse (1875) Spilota acutisterna Fairmaire (1878) P. lucidula,. 2 ohbayashii Nomura (1965). 2 pubicollis.. : Anomala acutisterna Fairmaire, Ann. Soc. Ent. Fr., 1877: 106. : 3 ( 1 ). :,,. : Reitter (1903), Kim (1996a, 2001b).

: : 79 26. Proagopertha lucidula (Faldermann), 1935: 380 ( 3-26, 11-26, 18-26) Anomala lucidula Faldermann, Mém. Acad. St. Petersb., II: 380, 1835. Phyllopertha lucidula: Murayama, 1935: 5; Miwa and Chûjô, 1939: 45; Sawada, 1941: 45; Cho, 1957: 293. Proagopertha lucidula: Stebnicka, 1980: 273; Kim et al., 1991e: 179; 2000: 132; ESK and KSAE, 1994: 153; Kim, 1996a: 106 ( ); 2001b: 94. Phyllopertha pubicollis (Waterhouse), Trans. Ent. Soc. Lond., 1875: 111 (Anomala): ( ): Okamoto, 1924: 177; Masaki, 1936: 261; Miwa and Chûjô, 1939: 45; Sawada, 1941: 44; Cho, 1957: 293; 1963: 219 ( ); Cho et al., 1968: 265; KZS, 1968: 135. Proagopertha pubicollis ( ): KSPP, 1972: 208; Kim et al., 1987c: 506; 1991b: 68 ( ); 1991d: 192; Park et al., 1993: 178; Park and Kim, 1993: 114. : (KZS, 1968). :.,..,.,,...,. (inner spine). Ad:.... 2. BL: 8 12 mm, BW: 4.5 7 mm. : Zhang, 1984: 60. :,. :,,,. : NK-HB, HN,, PB-,, SK-GW,,,,,, GS,,,,,,,,,,,,,,,,,,,, CBN,,,,,,,,,,, GBN,,,,, JBN,, JB-GN-JN. (236ex): [SWU] : 1 ( : 1933, Cho); 1 ( : 1933, Cho). : 1ex ( : 5.vi.1979); 1ex ( : 6.v.1984). : 1 ( : 1934, Cho); 2 ( : 1934, Cho); 3 ( : 1934, Cho); 2 ( : 1933, Cho); 50ex (,,,, : iv x.1958 2000). : 24ex (,,,,,, : iv vi.1984 1998). : 1ex ( : 26.v.1996). : 1ex ( : 26.v.1993). [NAAS] : 57ex (, : iii vi.1922 1989). : 2ex ( : 30.vi.1995).

80 : 5ex (, : iv v.1976 1989). : 1ex ( : 18.iv.1985). : 1ex (vi.19?). [SNU] : 1ex ( : 27.iv.1933). : 9ex ( 3, : iv viii.1955 1977). [KU] : 13ex (,, : iv vi.1957 1991). : 1ex ( : 20.iv.1983). [EWU] : 57ex (,,, : iv x.1959 1993). : 1ex ( : 14.x.1971). [ ] : 2 [ : 23.vi.1987 (5, 20.ix.1987 ), Kim]. : 219, 3.20. 10.17. (iii- 2, iv- 106, v- 67, vi- 36, viii- 2, ix- 2, x- 2ex). Genus Phyllopertha Stephens, 1830: 223..,,. (protibial inner spine) 2. 3 Ph. intermixta Blitopertha pallidipennis. : Scarabaeus horticola Linné, 1758. (Westwood, 1838 ). : 25 (Loebl and Smetana, 2006) ( 2 ). :. : Reitter (1903), Sawada (1938, 1941), Kim (1996a). 1.,,,.., diversa,.., horticola 27. Phyllopertha diversa Waterhouse, 1875: 106 ( 3-27, 11-27, 12-27, 18-27) Phyllopertha diversa Waterhouse, Trans. Ent. Soc. Lond.: 106, 1875: Heyden, 1887: 252; Okamoto, 1924: 178; Morita, 1936: 861; Doi, 1938: 97; Miwa and Chûjô, 1939: 49; Sawada, 1941: 48; Cho, 1957: 293;

: : 81 1963: 219; KZS, 1968: 135 ( ); Cho et al., 1968: 265; Hyun and Woo, 1970: 194; Stebnicka, 1980: 208; Kim et al., 1991b: 68; 1991d: 192; 1991e: 179; 1992c: 154; 1996b: 128; 2000: 132; 2002: 120; ESK and KSAE, 1994: 153; Kim, 1996a: 108; 2001b: 95; Lie et al., 1997: 36. Phyllopertha diversa koreana Ohaus, Stett. Ent. Zeit., 76: 325, 1915b: Masaki, 1936: 252 (coreana); Miwa and Chûjô, 1939: 49; Sawada, 1941: 48; KZS, 1968: 135; Kim, 1996a: 108 (syn. nov.). : (Cho, 1963), (KZS, 1968; KSPP, 1972). : (Nagasaki).,,,,..,.,,.,.. :. :. Ad:, 1.7.. 1.3,.,. BL: 7.5 9 mm, BW: 4 4.5 mm. : Zhang, 1984: 59. :,,. :,,,. : NK-HN, SK- 80 100 700. (557ex): [SWU] : 28ex (,,,,, : iv vi.1974 1993). : 140ex [,,,,,,, (5 ),,,,, : iv x.1935 2001]. : 12ex (,,,, : v.1987 1996). : 1ex ( : 8.vi. 1974). : 21ex (, : v.1993, 1994); 2ex ( : v.1977 1989). [NAAS] : 1ex ( : 13.vi.1973); 2ex ( : 30.v.1974). : 1ex ( : 11.vi.1996); 6ex ( : v vi.1924 1987). : 17ex ( : v.1998). : 12ex ( : v.1976). : 3ex ( : v.1986). : 1ex (29.v.1922). [SNU] : 21ex (,,,,,, : iv vi.1968 1992). : 1ex ( : 10.vi.1992). : 2ex ( : v.1990). [KU] : 123ex (,,, : vi.1974 1992). : 70ex (,,,,,, : v viii.1958 1985). : 10ex (,,,,, : 1974 1988). : 2ex ( : vi.1974 1981). : 14ex (,,, : v vii.1922 1988). : 7ex (v.1978). [EWU] : 3ex ( : vi.1979). : 30ex (,,,,,,, : v vi.1956 1987). : 1ex ( : 20.v.1988). : 1ex (7.v.1981). [HSU] : 23ex ( : v.1994 1997). : 518, 4.2. 10.23. (iv- 13, v- 294, vi- 207, vii- 1, viii- 1, x- 1ex).

82 : Ohaus (1915b) (Chemulpo) P. d. koreana. koreana,., 400 24 6%. 28. Phyllopertha horticola (Linné), 1758: 351 Scarabaeus horticola Linné, Syst. Nat., Ed. 10, I, 2: 351, 1758. Phyllopertha horticola: Stebnicka, 1980: 273; ESK and KSAE, 1994: 153; Kim, 1996a: 108 ( ); 2001b: 96.,.,..,,.,. 2.. BL: 9 11 mm, BW: 5 mm. : Medvedev, 1952: 67; Zhang, 1984: 59. :,,. :. : NK-Sunan and Sǒkam-Čǒsudži, PN (6, 21.v.1965, Morczekowski and Riedel: Stebnicka, 1980.). (3ex): [SWU] : 1 2 [ : 19.vii.1934, Cho ( )]. Genus Mimela Kirby, 1823: 92. Rhombonyx Hope, 1837 ( : holosericea Fabricius, 1787). Paracrucis Newman, 1839 ( : cyanipes Newman, 1839). Amblomala Reitter, 1903 ( : aurata Fabricius, 1801). Eriomela Reitter, 1903 ( : pomacea Bates, 1890).

: : 83 Paramimela Ohaus, 1915 ( : circumcincta Hope, 1842). Trimela Ohaus, 1924 ( : macassara Heller, 1896). :. Mimela Reitter (1903) Ohaus (1915, 1943). Machatschke (1952). 13 6 3 4. 2. : Mimela chinensis Kirby, 1825. Trans. Linn. Soc. Lond. 14: 101. : 162 (Loebl and smetana, 2006) ( 6 ). :. : Heyden (1887), Reitter (1903), Ohaus (1915a, 1943), Murayama (1938a), Machatschke (1952), Kim (1996a). 1. (Mimela s. str.).., 2 (subg. Paramimela).,., 3 2..,.., 1 fusania..,., splendens 3..,.., 1, 3 4 holosericea.,. 4 4 4.,.,. 1.

84 testaceipes, 5 5..,. Ad: costata coreana., pekinensis coreana 29. Mimela fusania Bates, 1888: 375 ( 3-28, 12-28, 18-28) Mimela fusania Bates, Proc. Z. Soc. Lond., 26: 375, 1888: Ohaus, 1915a: 95; Niijima and Kinoshita, 1923: 119; Maruta, 1929: 370; Masaki, 1936: 260; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 40; Cho, 1957: 294; 1969: 681 ( ); Kim and Kim, 1974a: 229; 1975: 247; Kim and Nam, 1982a: 156; 1982b: 276; 1984c: 87; 1985: 105; Kim, 1978: 361; Stebnicka, 1980: 209; Nam and Kim, 1983: 129; Yoon et al., 1990a: 140; Kim et al., 1987c: 506; 1991b: 68; 1991d: 192; 1991e: 179; 2001: 132; 2005: 79; Park et al., 1992: 178; Park and Kim, 1993: 113; ESK and KSAE, 1994: 153; Kim, 1996b: 237; 2001b: 98. Mimela chinensis Kirby, 1823 ( ): Stebnicka, 1980: 209; ESK and KSAE, 1994: 153. : (KSPP, 1972), (Yoon et al., 1990a); chinensis= (ESK and KSAE, 1994). : Fusan (= )..,.,..., 1.. Ad:..,.. BL: 13 17 mm, BW: 7.5 10 mm. :,,.

: : 85 :,,. : NK-HN, PN, SK- (20 ) 80 140 1,000. (732ex): [SWU] : 1ex ( : 6.vi.1974). : 77ex [,,, (6 ),,,,, : iv viii.1959 1993]. : 5ex (, : v vi.1974 1994). : 2ex ( : vi.1985). : 1ex ( : 22.vi.1973). : 1ex ( : 2.v.1984). [NAAS] : 24ex (,,, : iv vii.1921 1986). : 3ex ( : iv.1929). : 2ex (, : v.1976, 1991). [SNU] : 43ex (,, : iv x.1958 1995). : 1ex ( : 29.v.1997). [KU] : 1ex ( : 13.v.1934); 361ex (,,,, : iv ix.1959 1992). : 2ex ( : v.1986). : 2ex (, : v, vi.1977, 1991). [EWU] : 3ex (, : v viii.1966 1989). : 193ex (,,,,,,,, : iv x.1956 1988). : 2ex ( : v.1972). : 2ex ( : iv, viii.1964, 1972). : 5ex (,, : iv x.1971 1985). : 576, 4.21. 10.5. (iv- 75, v- 443, vi- 37, vii- 4, viii- 3, ix- 12, x- 2ex). : Nakayama and Okamoto, 1940. : 1970 (Kim, 1998c: 351). 30. Mimela splendens (Gyllenhal), 1817: 110 ( 3-29, 12-29, 18-29) Melolontha splendens Gyllenhal, in Schönh., Syst. Ins., I.3, Append.: 110, 1817. Mimela splendens: Bates, 1888: 375; Maruta, 1929: 370; Yoshino, 1935a: 14; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 47; Kim, 1956: 9; Cho, 1957: 294; 1963: 219 ( ); 1969: 680; Cho et al., 1968: 265; Ku, 1963: 28; Hyun and Woo, 1969: 193; Kim and Kim, 1975: 248; Kim and Nam, 1977: 131; 1982a: 156; 1982b: 276; 1984a: 103; 1984b: 328; Kim, 1978: 362; Yoon and Nam, 1979: 149; 1990b: 111; Kim, 1981: 345; 1995b: 140; 1996b: 237; 1997: 231; 2001b: 99; Kim et al., 1987b: 224; 1987c: 506; 1991b: 68; 1991d: 192; 1991e: 179; 1994b: 111; 1996b: 128; 2001: 132; 2004: 117; 2005: 79; Stebnicka, 1980: 274; Nam and Kim, 1983: 129; Park and Cho, 1986: 128; 1992: 178; Park and Park, 1987: 673; Park and Kim, 1993: 113; ESK and KSAE, 1994: 153; Lie et al., 1997: 45. Mimela splendens corusca Heyden, Horae Soc. Ent. Ross., 21: 253, 1887: Miwa and Chûjô, 1939: 47; Cho, 1957: 294. Mimela splendens cyanicollis Ohaus, Stett. Ent. Zeit., 76: 96, 1915b (Chemulpo): Niijima and Kinoshita, 1923: 119; Miwa and Chûjô, 1939: 47; Cho, 1957: 294. Anomala difficilis Waterhouse, Trans. Ent. Soc. Lond., 1875: 111: Heyden, 1887: 252. Mimela difficilis: Reitter, 1903: 53; Kim et al., 1992b: 154; Park et al., 1993: 178. Mimela lucidula Hope, Trans. Ent. Soc. Lond.: 113, 1835: Niijima and Kinoshita, 1923: 117; Okamoto, 1924: 177.

86 Mimela lucidula Hope var. gaschkewitschi Motschulsky, Etud. Entom., 6: 32, 1857: Heyden, 1887: 253. Mimela lucidula Hope var. coruscus Heyden, Horae Soc. Ent. Ross., 21: 253, 1887. Mimela flavilabris (Waterhouse, 1875)( ): KSPP, 1972: 207; Kim and Chang, 1984: 169; ESK and KSAE 1994: 153. : splendens= (Gu, 1963); (KSPP, 1972); (Ku, 1973); difficilis= (Park et al., 1993).,..,,.,.,.. Ad:,. ( ),,.,. BL: 15 21 mm, BW: 9 11 mm. : Zhang, 1984: 56. :,,,,. :,,,. : NK-HN, PB, PN, WN, GW, SK- (20 ) 100 130 3,000. (1,787ex): [SWU] : 11ex (,,, : v viii.1974 1997); : 278ex [,,,,,,,,,, (4 ),,, : v ix.1957 1991]. : 91ex (,,,,, : vi ix.1960 1998); 2ex ( : 17.vi.2003); 2ex ( : 23.vi.2005). : 6ex ( : vi vii.1986 1989). : 38ex (,,,,, : v xi.1922 1993). : 8ex (vi viii.1983 1997). [NAAS] NK-1ex ( : 25.vi.1922). : 5ex (, : vi.1973 1996). : 58ex ( : i ix.1921 1996); 3ex (, : viii.1971, 1976). : 8ex (,, : vi vii.1960 1996). : 1ex ( : 3.vi.1979). : 17ex (,,, : vi viii.1924 1991). : 145ex (v vii.1922 1993). [SNU] : 1ex ( : 12.ix.1993); 1ex ( : 23.vi.1986). : 69ex [,,,, (3 ),,,,, : iv x.1957 1995]. : 12ex (,, : vi xi.1960 1993). : 120ex (,, : vi vii.1924 1996). : 18ex (vii.1965 1990). [KU] NK-3ex (,?: vi vii. 1917). : 1ex ( PB: 7.vi.1931). : 2ex ( : vii.1975); 2ex ( : vi.1984). : 264ex [,,,, (5 ),,,,, : v ix.1958 1992]. : 6ex (,, : v vii.1979 1984). : 72ex (,,,,, : iv viii.1960 1990). : 20ex (,,,, : v viii.1973 1993). : 8ex (vii ix.1959 1973).?: 8ex ( : vi.1988). [EWU] : 16ex (, : iv x.1970 1973). : 421ex (,,,,,,,,,, : iv x.1955 1999). : 21ex (,,,,

: : 87 : vi viii.1967 1971). : 9ex (,,, : vi x.1969 1983). : 11ex (v viii.1965 1987). [HSU] : 34ex ( : iv vi.1994 1997). : 1,490, 4.11. 11.28. (iv- 11, v- 59, vi- 1,016, vii- 327, viii- 37, ix- 30, x- 9, xi- 1ex). 31. Mimela holosericea (Fabricius), 1787: 21 ( 3-30, 12-30, 18-30) Melolontha holosericea Fabricius, Mant. Ins., I: 21, 1787. Rhombonyx holosericeus: Heyden, 1887: 252; Stebnicka, 1980: 275 (holosericea). Anomala holosericea: Niijima and Kinoshita, 1923: 86; Cho, 1934: 75; Kim and Kim, 1975: 246; Kim and Nam, 1981: 125; 1982a: 156; 1984a: 103; 1984b: 328; Kim, 1978: 357. Mimela holosericea: Mochizuki and Tsunekawa, 1936: 211; 1937: 91; Miwa and Chûjô, 1939: 46; Nagaoka, 1940: 474; Mori and Cho, 1940: 13; 1947: 65 ( ); 1957: 293; 1969: 680 ( ); Cho et al., 1967: 297; Kim, 1960: 27; Won and Choi, 1968: 379; Kim and Kim, 1974a: 229; Kim and Nam, 1984b: 328; Kim et al., 1987a: 104; 1991b: 68; 1991d: 192; 1991e: 179; 1994a: 147; 1996b: 128; 1998: 170; 1999: 130; 2000: 132; 2002: 120; Kim, 1992: 106; 1993: 65; 1995a: 165; 1996b: 238; 1996c: 174; 2001: 100; ESK and KSAE, 1994: 153; Lie et al., 1997: 44..,...... 4 1. Ad:.,,. BL: 18 20 mm, BW: 6 9 mm. : Zhang, 1984: 55. :,,,,,. :,.. : NK-HB,, YG,,, HN Hosan (=, ),, PB,,,, GW,, NK?-Shoko, Sounando, Kazando, Koshou, SK- GW,,,,,,,,,, GS,,,,,,,, CB,, GB,,, JN,,. (124ex): [SWU] : 4ex ( : vi viii.1995 2000); 27ex ( : vi viii.1958 1997); 3ex

88 ( : vi vii.1985). : 1ex ( : 11.vi.1990); 2ex ( : vi, viii.1991, 1993). : 3ex (, : vi viii.1989 1994). : 13ex (,, : vi viii.1986 1998); 12ex ( : 29.vi 3.vii.1998); 9ex ( : 30.vi.1998). : 1ex ( : 14.viii.1992). [NAAS] NK- : 1ex [Hosan (= ): 8.vii.193?]; 1ex ( : 24.vii.1925). [SNU] : 1ex ( : 21.vii.1974); 1ex ( : 26.viii.1996). : 1ex ( : 20.vi.1992). : 1ex ( : 20.v.1984). : 4ex ( : vi vii.1976 1993). [KU] NK: : 2ex [Ketzanchin (= ): vii viii.1933, 1934]; 3ex ( : vii.1932); 5ex ( : vii viii. 1930 1937). : 6ex (,,,, : vi viii.1979 1994).?: 2ex (Shoko: 16.viii.1931); 2ex (Sounando: vii.1937); 2ex (Kazando: vii, viii.1937); 2ex (Koshou: viii.1936). [EWU] : 5ex ( : 30.vii.1974); 3ex ( : 9.vii.1995). : 2ex ( : vi.1984). [HMNH-NK] YG: 2ex ( : 27.vii.1975). : 1ex ( : 13 4.ix.1980); 2ex ( : 9 11.vii.1977); (17 21.vi.1988). : 124 9 1 6 8. 32. Mimela testaceipes (Motschulsky), 1860: 14 ( 4-31, 12-31, 18-31) Anomala testaceipes Motschulsky, Etud. Ent., 9: 14, 1860: Heyden, 1887: 252; Bates, 1888: 374; Niijima and Kinoshita, 1923: 87; Maruta, 1929: 369; Mochizuki, 1936: 211; Morita, 1936: 860; Masaki, 1936: 250; Doi, 1937: 63; Cho, 1947: 65: 1957: 293; Kim and Kim, 1976: 102; Kim and Nam, 1981: 125; 1982a: 156; Kim, 1978: 354; Shin, 1978: 143; Kim, 1981: 344; 1995a: 165; 1995b: 140; 2001: 101; Kim and Chang, 1982a: 145; Park and Cho, 1986: 128. Mimela (Rhombonyx) testaceipes: Reitter, 1903: 56. Paramimela testaceipes: Ohaus, 1915a: 91. Mimela testaceipes: Miwa and Chûjô, 1939: 46; Machatschke, 1952: 362; Hyun and Woo, 1969: 193; Kim et al., 1974b: 108; Nam and Kim, 1983: 129; Ku, 1963: 28 ( ); Kim et al., 1987a: 104; 1987b: 224; 1987c: 506; 1991e. 179; 1991a: 232; 1991b: 68; 1991d: 192; 1992b: 154; 1994a: 147; 1994b: 111; 1996a: 59; 1996b: 128; 1998: 170; 1999: 130; 2000: 132; 2002: 120; 2003: 126; 2005: 79; Yoon et al., 1990a: 140; Park et al., 1993; 178; Park and Kim, 1993: 114; Kim, 1992: 106; 1993: 65; 1995a: 165; 1995b: 140; 1996b: 238; 1996c: 174; 1997: 231; 2000b: 101; ESK and KSAE, 1994: 153; Lie et al., 1997: 43. Rhombonyx testaceipes: Stebnicka, 1980: 209. Anomala testaceipes ochroptera Ohaus, Stett. Ent. Zeit., 76: 91, 1915: Niijima and Kinoshita, 1923: 88; Doi, 1935: 3; Kamijo, 1936: 95; Morita, 1936: 860. Rhombonyx ussuriensis Medvedev, 1951: Stebnicka, 1980: 209. Rhombonyx testaceipes ussuriensis: Kalinina, 1989: 411. : (Hyun and Woo, 1969 6). :.

: : 89,..,.,. 4., 1.. Ad: 2/3, ( ),... BL: 14 20 mm, BL: 6 9 mm. : Murayama, 1931: 56. :,,,. :,,,,. : NK-HB, YG,,, HN, PB,,,,, PY,, WN, GW,,? Tschon-Bon-San, Nyompjang, Kuson (Tschondtsin), Kekki (Vekki), SK- 140 180 1,500,. (1,117ex): [SWU] : 60ex (,,,,,,,,,,,, : vii viii.1976 2001); 7ex ( / : 10/20.vii.2009). : 184ex [,,,,,,,,, (9 ),,,,, : vi x. 1959 2000). : 25ex (,,,,,,,, : vi viii.1960 1999); 1ex ( : 27.vii.2008). : 111ex (,,,,,,,,,,,,, : vi ix.1981 2000). : 29ex (,,,,,,, : 1983 1993). : 7ex (vii.1985). [NAAS] NK-1ex ( : 25.vii.1924); 2ex ( : vii.1921, 1924). : 43ex (,,, : vii.1995, 1998); (viii.1987). : 86ex (,,,,,, : v ix.1921 1998). : 17ex (, : viii x.1996 1998). : 10ex (,,,, : vi viii.1982 1997). : 10ex (,,,, : v xi.1922 1996). : 2ex (7.vi.1986). [SNU] : 3ex (,, : vii, viii.1974, 1995, 1996). : 31ex (,,,, : v xi.1963 1997). : 5ex (,, : vii.1983 1997). ; 5ex ( : vi.1994). : 101ex (, : vi viii.1926 1997).?: 2ex ( : viii.1993). [KU] NK-2ex ( : 5.vii.1916). : 10ex (,, : vii.1958 1981). : 82ex (,,,,, : vii viii.1934 1984). : 42ex (,, : vii ix.1957 1988). : 35ex (,,,,,, : vi viii.1960 1990). : 62ex (,,, : vii viii.1974 1985).?: 2ex (Eusenrei: 27.vii.1933). [EWU] : 22ex (,,, : vii viii.1970 1983). : 43ex (,,,,,, : v x.1956 1987). : 7ex (,, : vii.1960 1973). : 10ex (,,,, : vii viii.1956 1986). : 5ex (,, : vii viii.1959 1977). [HSU] : 2ex ( : vi.1996, 1997). [HMNH-NK] : 1ex ( : 10.vii.1982). : 5ex ( : 13,

90 14.vii.1982). : 1ex ( : 4 6.viii.1975); 37ex (9 11.vii.1977); 5ex (22 25.vii.1982). : 1,024, 5.2. 11.20. (v- 10, vi- 147, vii- 555, viii- 275, ix- 18, x- 6, xi- 3ex). : Murayama, 1936; Nakayama et Okamoto, 1940; Torikura, 1991 ( ); Kim and Kang, 1993: 415. 33. Mimela pekinensis coreana Machatschke, 1952: 367 Anomala pekinensis Heyden, Deut. Ent. Zeit., 30: 291, 1886. Mimela pekinensis coreana Machatschke, Beitr. Entom., Berlin, 2(4/5): 367, 1952: Kim, 1996b: 240 ( ); 2001b: 103. Mimela pekinensis koreana: Stebnicka, 1980: 209; ESK and KSAE, 1994: 153. : (ESK and KSAE, 1994). : (=, Che-mul-po Je-mul-po ).,..,. Machatschke (1952),,. coreana. :,. :?. : ( : 1, : 1 4, : Deut. Entom. Inst., Berlin). :. :. 34. Mimela costata (Hope), 1839: 73 Euchlora costata Hope, Proc. Zool. Soc. Lond., VII: 73, 1839. Anomala costata: Nam and Kim, 1983: 128. Mimela costata: Kim and Chang, 1987a: 104; Kim, 1992: 106; 1996b: 240; 2001b: 104; ESK and KSAE, 1994: 153 ( ).

: : 91,,. Machatschke (1952),,,. BL: 16 21 mm. :,. :?. : (Kim, 1996b; 2001b). : [HMNH-NK] Korea, 1ex, Reitter, 1893. :, Machatschke (1952) Ohaus. Hungary 1 Korea, Reitter, 1893. Reitter, Anomala. 1893 1898. Reitter.. Mimela (= ) 1. Mimela concolor Blanchard, 1851 Mimela concolor Blanchard, Cat. Coll. Ent. Mus. Hist. Nat. Paris, Coleopt. I (2): 196, 1851: Stebnicka, 1980: 209 ( ); ESK and KSAE, 1994: 153 ( ) ( ),. 2. Mimela coreana Niijima and Kinoshita? Mimela coreana Niijima and Kinoshita: Mochizuki and Tsunekawa, 1937: 91, Niijima and Kinoshita (1923) Miridiva coreana,. 3. Mimela flavilabris (Waterhouse), 1875 Anomala flavilabris Waterhouse, Trans. Ent. Soc. Lond.: 110, 1875: Mimela fl.-: Kim and Chang, 1987:

92 169 ( ); KSPP, 1972: 207 ( ); ESK and KSAE, 1994: 153 (, ),. 4. Mimela testaceoviridis Blanchard, 1850 Mimela testaceoviridis Blanchard, Cat. Coll. Ent. Mus. Hist. Nat. Paris, Col. I (2): 197, 1850: Niijima and Kinoshita, 1927: 57; Miwa and Chûjô, 1939: 48; Cho, 1957: 294; KZS, 1968: 135 ( ), KSPP, 1972: 208; ESK and KSAE, 1994: 135 ( ) M. testaceipes. Genus Callistethus Blanchard, 1851: 198. Anomala Spilota. Machatschke (1955) 32 S. plagiicollis praefica 2, 1. : Callistethus consularis Blanchard, 1851. : 2 (Machatschke, 1955) ( 1 ). :,. : Machatschke (1955), Kim (1996b, 2001a, b). 35. Callistethus plagiicollis (Fairmaire), 1886: 329 ( 4-32, 12-32, 18-32) Anomala plagiicollis Fairmaire, Ann. Soc. Ent. Fr., (6)VI: 329, 1886: Murayama, 1935: 5; 1937a: 35; 1937b: 14; Mori and Cho, 1940: 13; Cho, 1947: 65; 1957: 292. Anomala (Spilota) plagiicollis: Niijima and Kinoshita, 1923: 116; 1927: 41; Miwa and Chûjô, 1939: 42. Anomala (Anomala) plagiicollis: Miwa and Chûjô, 1939: 38. Anomala (Spilota) plagiicollis impicta: Miwa and Chûjô, 1939: 42; Cho, 1957: 292. Spilota plagiicollis: Reitter, 1903: 51; Stebnicka, 1980: 273; Kim et al., 1991b: 68 ( ); 1991c: 154; 1991d: 193; 1991e: 179; 1992b: 154; 1994b: 111; 1996a: 59; 1996b: 128; 1997: 231; 1998: 170; 1999:

: : 93 130; 2002: 120; 2003: 126; 2004: 117; Kim, 1992: 106; 1993: 65; 1995a: 165; 1995b: 140; 1996a: 109; 1996c: 174; 2001b: 105; Park and Kim, 1993: 114; ESK and KSAE, 1994: 153 (plagicollis). Callistethus plagiicollis: Lie et al., 1997: 35. Anomala (Paraspilota) impicta Bates, Proc. Zool. Soc. Lond., 1888: 374 (Corea). : (Cho, 1947); (KZS, 1968). :,..,.,.,. ( ).,. Ad:,.,.. BL: 12 18 mm, BW: 5.5 9 mm. :,,. :,,,. : NK-YG, HN, PB, GW, SK (1990 )- ( ),,,,,,, SK (1990 )- 100 130 1,200,. (1,057ex): [SWU] : 93ex (,,,,,,,,,,,,,, : v viii.1962 2001). : 55ex [,,,,, (6 ),,,, : iv viii.1957 1996]. : 47ex (,,,,,,, : v viii.1960 1998). : 104ex (,,,,,,,,,,,, : vi ix.1975 1998); 227ex ( : 29.vi 3.vii.1998); 3ex ( : 5, 6.vi.2008); 1ex ( : 19.vii.2001). : 22ex (,,,,,, : v viii.1976 2001). : 2ex (vii.1983, 1997). [NAAS] NK-3ex ( : vii, viii.1924). : 15ex (,,, : vi viii.1923 1997). : 12ex (,, : vi viii.1993 1997). : 1ex ( : 26.vi.1991). : 52ex (,,,, : vi ix.1983 1997). : 6ex (,, : vi vii.1922 1995). : 2ex (vi, vii.1976, 1958 1994). : 28ex [,, (4 ),, : v xii. 1958 1996). : 5ex (,, : vii.1957 1988). : 25ex (,,, : vi viii.1960 1990). : 30ex (,, : vi viii.1976 1993). : 2ex (2.viii.1959). [EWU] : 10ex (,,, : vi vii.1973 1995). : 29ex (,, : vi viii. 1959 1984). : 1ex ( : 22.vii.1968). : 3ex ( : vii.1983); 1ex ( : 10.viii.1959). : 1ex (31.v.1982). [HSU] : 64ex ( : iv vi.1995 1997). [HMNH-NK] : 1ex ( : 18.vii. 1977). : 3ex ( : 3.vii.1991). : 1ex ( : 4.viii.1975); 5ex (9 12.vii.1977); 1ex (22.vii.1982);

94 1ex (17.vi.1988); 8ex (9 12.vi.1991). : 716, 5.9. 12.26. (v- 16, vi- 279, vii- 338, viii- 80, ix- 9, x- 3, xi- 3, xii- 1ex). : Murayama (1935b) ( : 5, 5 ).. Genus Blitopertha Reitter, 1903: 85. Spilota Anomala, Exomala., Blitopertha. : Melolontha lineata Fabricius, 1798. Suppl. Entom. Syst.: 525. : 30 (?), 13 (Reitter, 1903: 3 ) (2 3 ). :,. : Reitter (1903), Sawada (1938, 1941), Kim (1996a, 2001a, b). 1.. (subg. Blitopertha) 11 mm.,.,. conspurcata.. (subg. Exomala) 10 mm 13 mm.,,. 2 2.,., pallidipennis. 2 3., orientalis

: : 95 36. Blitopertha conspurcata (Harold), 1878: 71 ( 4-33, 12-33, 18-33) Phyllopertha conspurcata Harold, Deut. Ent. Zeit., 22: 71, 1878 ( conspercata ): Heyden, 1887: 252; Bates, 1888: 373; Lewis, 1895: 400; Niijima and Kinoshita, 1923: 120; Cho, 1934: 75; 1969: 679; Cho et al., 1967: 198 ( ); Haku, 1937: 121; Masaki, 1936: 261; Doi, 1937: 62; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 44; Sawada, 1941: 54; Kim and Kim, 1974a: 229; 1975: 246; Kim and Nam, 1984a: 103; 1984c: 87. Blitopertha conspurcata ( ): Kim and Kim, 1971: 160; Kim and Nam, 1982a: 157; 1982b: 277; 1984b: 328; 1985: 105; Kim, 1978: 374; Yoon and Nam, 1979: 149; 1985: 160; Stebnicka, 1980: 279; Kim, 1981: 345; 1992: 106; 1993: 65; 1995a: 165; 1996a: 110; 2001: 107; Kim et al., 1987c: 506; 1991b: 68; 1991d: 192; 1991e: 179; 1994a: 147; 1998: 170; 2000: 132; 2001: 132; 2005: 79; 2006: 37; Park et al., 1993: 177; Park and Kim, 1993: 113; ESK and KSAE, 1994: 152; Kim, 1996a: 110. :.,.,.,.,,... Ad:, ( ),., 2. BL: 8 11 mm, BW: 4.5 5.5 mm. : Medvedev, 1952: 70; Zhang, 1984: 58. :,,,. :,,,. : NK-HB,, YG,, HN,, PB,,, PY,, PN, WN, GW-,, GG, SK- (20 ) 100 140 2,000. (1,464ex): [SWU] : 49ex (,,,,, : v vii.1980 1998). : 236ex [,,,,,, (6 ),,,, : iv x.1963 1997]; 1ex ( : 5.vi.2002). : 30ex (,,,,, : iv vi.1963 1997). : 11ex (,,, : v vi.1978 1998). : 26ex (,,,, : v vi.1987 1994). [NAAS] : 11ex (,, : v vi.1981 1993). : 19ex ( : v vi.1922 1993). : 10ex (, : v vi.1979 1986). : 14ex ( : viii.1983). [SNU] : 141ex (,,,,,, : v vi.1968 1994). : 5ex (,, : vi v.1984 1992). : 1ex ( : 13.vii.1995). : 6ex (,, : v viii.1990 1999). [KU] NK- : 2ex [Ketzanchin (=

96 ): vi.1934]. 3ex [, Mt. Ganbozan (= ): 14, 18.vii.1932]. : 35ex (,,, : v, vi.1974 1979). : 339ex [,,,,,,,,, (7 ), : v vi.1931 1992]. : 82ex (,,,,,, : 1974 1987). : 11ex (,,,, : v vi.1974 1996). : 46ex (,,, : v vi.1972 1993). [EWU] : 6ex (, : v vi.1972, 1974); 5ex (,, : v vi.1979 1995). : 362ex (,,,,,,,,,,,, : iv x.1956 1993). : 1ex ( : 20.v.1988). : 1ex ( : 17.v.1987). : 2ex ( : v.1966). [HMNH-NK] : 9ex ( : 24 28.v.1991). : 1,358, 4.28. 10.3. (iv- 5, v- 670, vi- 726, vii- 42, viii- 9, ix- 4, x- 1ex). 37. Blitopertha pallidipennis Reitter, 1903: 89 ( 4-34, 12-34, 18-34) Blitopertha (Blitopertha) pallidipennis Reitter, Ver. Nat. Ver. Brünn, Best.-Tab., 51: 89, 1903. Phyllopertha pallidipennis ( nigrata ): Niijima and Kinoshita, 1923: 212; Cho, 1934: 75; 1947: 65 ( ; var. nigrata= ); 1957: 293; 1969: 679; Murayama, 1935: 197; Masaki, 1936: 261; Haku, 1937: 121; Mochizuki, 1936: 211; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 44; Mori and Cho, 1940: 13; Sawada, 1941: 49; Hyun and Woo, 1970: 80; Kim and Kim, 1974b: 108; 1975: 247; 1976: 102; Kim and Nam, 1978: 134; 1981: 125; 1982a: 156; 1982b: 276; 1984a: 103; 1984b: 328; 1984c: 87; Yoon et al., 1978: 83; 1979: 149; 1990b: 111; Kim, 1981: 345; Park and Cho, 1986: 128. Blitopertha pallidipennis: Stebnicka, 1980: 249; Kim et al., 1987c: 506; 1991b: 68; 1991c: 154; 1991d: 192; 1991e: 179; 1992: 154; 1994a: 147; 1996a: 59; 1996b: 128; 1998: 170; 1999: 130; 2000: 132; 2001: 132; 2002: 120; 2003: 126; 2005: 79; Kim, 1992: 106; 1993: 65; 1995a: 165; 1995b: 140; 1996a: 111; 1996c: 174; 1997: 231; 2001: 108; Park et al., 1993: 177; Park and Kim, 1993: 113; ESK and KSAE, 1994: 152. : (Ku, 1973). : Ph. Pallidipennis-,,. Ph. pallidipennis nigrata-,.,,. ( nigrata ).,,..,, 1,.. Ad:.,. BL:

: : 97 8 12.5 mm, BW: 4.5 7 mm. : Medvedev, 1952: 69; Zhang, 1984: 58. :,,. :,,,. : NK-HB,,,, YG,,,, HN,,,,, PB,,, PY,, PN, WN, GW,, GG, SK- (40 ),,, 140 200 4,000. (2,686ex): [SWU] : 317ex [ ( ), : vi viii.1934 2000]. : 477ex ( 40 : iv x.1975 2001). : 92ex (,,,,,,,,,,,, : vi x.1960 1999); 1ex ( : 5.vii.2008). : 164ex (,,,,,,,,,,,,,,,, : vi viii.1979 2002). : 35ex (,,,,, : vi viii.1988 1994). : 1ex (17.viii.1991). [NAAS] NK-1ex ( : 18.vii.1923); 1ex ( : 7.vii.1924). : 1ex ( : 25.vi.1922). : 1ex ( : viii.1987). : 24ex (,, : vi xi.1924 1993). :(,, : vii, viii.1974 1983). : 4ex (,,, : vi, vii.1922, 1986 1988).?: 1ex (Yotori: 8.iv.1931). [SNU] : 21ex (,,,, : vi vii.1974 1997). : 85ex [,,, (3 ),,, : iv ix.1959 1997]. : 32ex (,, : vi vii.1983 1992). : 4ex (, : vii.1995, viii.1996). : 18ex (,,, : v viii.1968 1997). [KU] NK-1ex ( : 15.vii.1926); 2ex ( : vii.1930). : 145ex (,,,,,, : iv viii.1931 1995). : 331ex [,,,,, (7 ),,,,, : iv ix.1931 1996]. : 222ex (,,,, : v viii.1957 1983). : 290ex (,,,,,, : vi viii.1960 1994). : 18ex (,, : v viii.1974 1988). : 5ex (v viii. 1933 1974). [EWU] : 43ex (,,,,,, : iv x.1958 1984). : 303ex (,,,,,,, : v x.1955 1994). : 13ex (,,,, : vi vii.1969 1993). : 2ex ( : vii.1983). : 1ex ( : 3.viii.1974); 2ex ( : vi.1961). [HSU] : 13ex ( : 19.viii.1997). [HMNH-NK] : 1ex [ (1,150 m): 22.vii.1975]. : 1ex ( : 8.vii.1982). : 1ex ( : 18.vii.1982); 6ex (3 6.vii.1991); 3ex ( : 11 12.vii.1991). : 1,789, 4.8. 10.21. (iv- 19, v- 20, vi- 495, vii- 995, viii- 260, ix- 20, x- 9, xi- 1ex). : 1,218 ( 809+nigrata 409 ) ( )= : =18:1,200 (nigrata 9: 400). 1.5% (nigrata 2.25%). (B. orientalis).

98 38. Blitopertha orientalis (Waterhouse), 1875: 108 ( 4-35, 12-35, 18-35) Phyllopertha orientalis Waterhouse, Trans. Ent. Soc. Lond., 1875: 108: Yoshino, 1935a: 14; 1935b: 3; Ku, 1963: 28. Anomala orientalis: Heyden, 1887: 252; Bates, 1888: 374. Blitopertha (Exomala) orientalis: Reitter, 1903: 89. Blitopertha orientalis: KSPP, 1972: 206 (Britopertha, ); Stebnicka, 1980: 206; Kim et al., 1991e: 179; 1996a: 112; 1996b: 128; 1999: 130; 2000: 132; 2002: 120; 2005: 79; 2006: 37; Kim, 1996c: 174; 1997: 231; 2001b: 110; ESK and KSAE, 1994: 152. : (Kawachi, Nagasaki, Hakodadi)., 2 3.,.,. Ad:,. BL: 8 13.5 mm, BW: 4.5 7.5 mm. :,,,,,. :,,,. : NK-HB Poro-čhŭn ( 20 km), HN, PB, SK- (, ) 80 100 600. (439ex): [SWU] : 33ex (,,,, : vi viii.1993 1998); 1ex ( : 4.vii.2009). : 162ex [,,,,,,,, (Seiryori = 10 ),,,,,,, : vi ix.1933 1999]; 19ex [ (,,,, ): vi viii.2006 2009]; 1ex ( : 5.vii.2009). : 10ex (,,,, : vi viii.1987 1998). : 39ex (,,,,,,,,,,, : vi viii.1987 2000); 1ex ( : 5.vi.2008); 1ex ( : 19.vii.2001). : 8ex (, : vii.1981, 1985). : 3ex (iii vii.1985 1995). [NAAS] NK-2ex ( : vii.1923, 1924). : 6ex (,, : vii viii.1923 1987). : 17ex (,, : iii xi.1967 1993). : 5ex (,,, : vi viii.1987 1996). : 15ex (, : vi.1986 1991). [SNU] : 6ex ( : vii.1974 1997). : 38ex (,,,,,, : v xi.1969 1997). : 12ex (, : vi vii.1983 1991). : 1ex ( : 13.vii.1995). : 6ex (,, : vi ix.1982 1995). [KU] : 6ex (,, : vi viii.1985, 1993 1995). : 13ex (,,,, : iv viii.1974 1991). : 1ex ( : 25.vi.1994). : 4ex (,, : vi vii.1984 1990). : 2ex (, : vii.1985, vi.1993). : 1ex (2.viii.1974). [EWU] : 21ex (,,,, : vi x.1956 1994). : 1ex ( : 26.ix.1966). : 1ex

: : 99 ( : 16.vi.1979). [HSU] : 3ex ( : 19.viii.1997). : 372, 3.20. 11.18. (iii- 1, iv- 3, v- 6, vi- 160, vii- 148, viii- 43, ix- 9ex. xi- 2ex). : (B. pallidipennis)... 1990 ( ).. Genus Anomala Samuelle, 1819: 191. Idiocnema Faldermann, 1835 ( : Idiocnema sulcipennis Faldermann). Psammoscapheus Motschulsky, 1835 ( : Psammoscapheus dilutus Motschulsky). Heteroplia Burmeister, 1844 ( : Melolontha elata Fabricius). Rhinoplia Burmeister, 1844 ( : Melolontha dorsalis Fabricius). Nongoma Péringuey, 1902 ( : Anomala calcarata Arrow). Chrysoplethisa Reitter, 1903 ( : Anomala octiescostata Burmeister). Dichomala Reitter, 1903 ( : Melolontha devota Rossi). Diplomala Reitter, 1903 ( : Anomala subvittata R.). Emphalena Reitter, 1903 ( : Anomala exoleta Faldermann). Euchronomala Reitter, 1903 ( : Euchlora albopilosa Hope). Euporochlora Reitter, 1903 ( : Melolontha viridis Fabricius). Euporomala Reitter, 1903 ( : Anomala sieversi Heyden). Hybalonomala Reitter, 1903 ( : Anomala bleusei Chobuat). Idiocnemina Reitter, 1903 ( : Anomala gracilenta Reitter). Orphnomala Reitter, 1903 ( : Anomala rufozonula Fairmaire). Paragematis Reitter, 1903 ( : Anomala melanopa Reitter). Anomalepta Casey, 1915 ( : Anomala semilivida LeConte). Anomalopus Casey, 1915 ( : Anomala rhizotrogoides Blanchard). Hemispilota Casey, 1915 ( : Anomala lucicola Fabricius). Lamoana Casey, 1915 ( : Anomala villosella Blanchard). Paranomala Casey, 1915 ( : Melolontha binotata Gyllenhal). Rhombonalia Casey, 1915 ( : Anomala cavifrons LeConte). Iliola Seminov and Medvedev in Medvedev, 1949 ( : Iliola pallens Semenov).

100.., 2 4.,. Anomalina 1,000 800 (Jameson et al., 2003). 3 46. 46 7, 20 7. : Melolontha frischii Fabricius, 1775. : 1,000 225 (Loebl and Smetana 2006) ( 13 ). :,. : Reitter (1903), Ohaus (1915a, 1915b), Kim (1997a, 1998a, 1998b). (Ad LPMB= : : ) 1. 2 6 2. 14 mm,.. Ad:,. LPMB=1:1.04:1.3 sieversi 17 mm,. Ad:, 3 3.,,.,. Ad:,. LPMB=1:1.1: 1.3 albopilosa. 2, 2 4 4.,.. Ad:,. LPMB=1:1.3: 1.2 japonica,. 5 5.,. Ad:. LPMB=1:1.1:1.13 mongolica

: : 101,. Ad:,. LPMB=1:1.1:1.0 cuprea 6. 1 3, 4 7. 14 18 mm 10 7. 4,. geniculata 3, 8 8.,,,,.. 2/3. Ad:,,. LPMB=1:0.6:0.9 chamaeleon, 9 9. 12 mm.. 2/3. Ad:,. LPMB=1:0.95:1.05 rufocuprea 13 mm,,.. Ad:., koreana 10...,,,, 11 2/3... 12 11.... Ad: 1. LPMB=1:1.1:1.2 luculenta... Ad:, ogloblini 12... Ad: ( ). LPMB=1: 1.2: 1.4 corpulenta 2/3.. Ad:.. ignicolor

102 39. ( ) Anomala sieversi Heyden, 1887: 266 ( 4-36, 12-36, 18-36) Anomala sieversi Heyden, Horae Soc. ent. Ross., 21: 266, 1887: Bates, 1888: 374; Lewis, 1895: 401; Niijima and Kinoshita, 1923: 92; Okamoto, 1924: 175; Kamijo, 1932: 21; Masaki, 1936: 260; Mochizuki and Tsunekawa, 1937: 93; Miwa and Chûjô, 1939: 40; Cho, 1957: 292; 1963: 219; 1969: 678; Cho et al., 1968: 264; Kim and Kim, 1972a: 84; 1974a: 229; 1975: 245; Kim and Nam, 1982a: 156; 1982b: 276; 1984b: 328; 1985: 105; Kim, 1978: 352; Yoon and Nam, 1979: 149; Yoon et al., 1990b: 111; Stebnicka, 1980: 276; Kim, 1981: 344; 1998a: 303; 2001b: 113; Kim et al., 1987c: 506; 1991d: 193; 1991e: 179; 1997: 231; 2000: 132; 2001b: 132; 2006: 37; Park et al., 1993: 178; Park and Kim, 1993: 113; ESK and KSAE, 1994: 152. Anomala (Euporomala) sieversi: Reitter, 1903: 63. Phyllopertha sieversi: Sawada, 1941: 55, 56. Anomala sieversi atrocoerulea Reitter, Ver. Nat. Ver. Brünn. Best-Tab., 51: 63, 1902 (Corea): Miwa and Chûjô, 1939: 40; Cho, 1957: 292. Anomala sieversi subpurpurea Reitter, Ver. Nat. Ver. Brünn. Best-Tab., 51: 63, 1902 (Corea): Miwa and Chûjô, 1939: 40; Cho, 1957: 292. Anomala octocostata Motschulsky, Etud. Ent. IX: 14, 1860 (Japan): Heyden, 1887: 252; Okamoto, 1924: 175; Yoshino, 1935a: 14. Anomala (Chrysoplethisa) octocostata: Reitter, 1903: 57. Anomala octiescostata Burmeister, Hand. Ent., IV(1): 243 (Phyllopertha) (Japan): Niijima and Kinoshita, 1923: 91; Miwa and Chûjô, 1939: 38; Cho, 1957: 220, 1963: 219; Cho et al., 1968: 264; Ku, 1963: 27; Stebnicka, 1980: 209. Phyllopertha octiescostata: Sawada, 1941: 55, 56. : sieversi= (Cho, 1963; Cho et al., 1968); (Kim, 1978 4); octiescostata= (Cho, 1963; Cho et al., 1968), ( ) (Gu, 1963 3), (Cho, 1969 14). : (, ). 12%.,.,,.,.,. 2 3,.. Ad:,. 2, LPMB=1: 1.04: 1.3. BL: 11 14 mm, BW: 6.5 8.5 mm. : Zhang, 1984: 54.

: : 103 :,. :,,,. : NK-HN, PY,, PN,,,, GW Čhŏnne (= ), GG, SK- ( 50 ) 120 150 2,000. (1,175ex): [SWU] : 16ex (, : iv vi.1984 1986). : 129ex [,,, (9 ): iv x.1941 1997]. : 60ex (,,,,,,,,,,,, : iv vi.1958 1997). : 2ex (, : v, vii.1987, 1988). : 5ex (,,, : v vii.1985 1993). : 8ex (iv v.1983 1997). [NAAS] : 1ex ( : v.1980). : 96ex (,,,,,, : iv viii.1922 1990). : 3ex (v, x.1992, 1996). [SNU] : 1ex ( : 12.ix.1983). : 101ex (,,,,,,,,,,, : iv x.1935 1994). : 6ex ( : v vi.1958 1963). : 1ex ( : 5.vi.1987). : 13ex (,,, : iv vii.1958 1997). : 4ex (14.v vii.1965 1992). [KU] : 9ex (,,,, : v vi.1970 1992). : 344ex [,,,,,, (9 ),, : iv xi.1934 1993]. : 7ex (,,,,, : v.1974 1987). : 1ex ( : vi.1989). : 8ex (,, : iv vi.1972 1991). : 5ex (v.1963 1978). [EWU]. 2ex (, : vi.1974, 1987). : 334ex (,,,,,,,, : iv ix.1958 1988). : 2ex (, : vi.1993, 1970). : 1ex ( : 5.vi.1983); 2ex (, : iv, vi.1966, 1969). : 3ex (, : v.1978 1985). : 11ex (v.1965 1988). : 752, 4.10. 11.12. (iv- 132, v- 455, vi- 137, vii- 6, viii- 7, ix- 7, x- 6, xi- 2ex). : Saito, 1928: 13, 1931: 473; Nakayama and Okamoto, 1940: 198. 40. Anomala albopilosa Hope, 1839: 70 ( 4-37, 12-37, 18-37) Anomala albopilosa Hope, Proc. Zool. Soc., Lond., 1839: 70: Okamoto, 1924: 176; Doi, 1935: 3; Yoshino, 1935: 14; Kamijo, 1936: 86; Mochizuki, 1936: 210; Mochizuki and Tsunekawa, 1937: 90; Masaki, 1936: 260; Miwa and Chûjô, 1939: 41; Cho, 1947: 65 ( ); 1955: 207; 1957: 293; 1963: 219; 1965: 196; 1969: 676; Cho et al., 1968: 265; Won and Choi, 1968: 379; Hyun and Woo, 1969: 192; Shin and Noh, 1970: 37; 1977: 88; Gu, 1973: 79; Kim and Kim, 1975: 244; Kim and Nam, 1982a: 156; Kim, 1978: 348; Yoon and Nam, 1978: 83; 1979: 149; 1986: 160; Yoon et al., 1990a: 140; Stebnicka, 1980: 209; Lee and Kwon, 1981: 154; Kim and Chang, 1982a: 145; 1982b: 173; 1983: 166; 1984: 169; Kim and Lee, 1989: 176; 1991b: 68; Kim et al., 2006: 36; Lee and Ko, 1988: 211; Park et al., 1993: 178; Kim, 1994: 214; 1998a: 303; 2001b: 114; ESK and KSAE, 1994: 152. : (Cho, 1965; Gu, 1973).

104,,....,.,.,.,.. Ad:,., LPMB=1:1.1:1.3. BL: 18 25 mm, BW: 9 13 mm. :,. :,,,,. : NK-YG, HN,, GW, SK- 20 70 1,000. (835ex): [SWU] : 2ex (, : vii, viii.1990, 1997). : 7ex (,,, : vii ix.1981 1996). : 5ex (, : vii viii.1981 1993). : 4ex ( : 25.vii.1922); 15ex (,,,,,, : vii viii.1964 1998). : 116ex (,,,,,,,,, : vii viii.1982 1998); ( : 21.vii.2002, K.D. Han); 1ex ( : 9.viii.2008). : 65ex (vi viii.1972 1999); 2ex ( : 16.viii.2009). : 2ex (30.vii. 1993). [NAAS] NK-1ex ( : 25.vii.1924). : 65ex (,, : vii viii.1991 1996). : 6ex (,, : v viii.1991 1997). : 61ex (,,,,,,,, : vi x.1924 1993). : 158ex (vi ix.1922 1993). [SNU] : 10ex (,,,, : vi ix.1960 1996). : 1ex ( : 29.vi.1994). : 2ex ( : viii.1996). : 1ex ( : 20.viii.1996); 23ex ( : vi viii.1959 1997). : 47ex (vii viii.1965 1997). [KU] : 2ex ( : vii.1964, 1979). : 1ex ( : 2.viii.1934). : 7ex ( : vii ix.1973 1979). : 2ex ( : viii.1928); 6ex ( : vii.1978). : 173ex (, ). :( : v viii.1959 1991). [EWU] :( ); 17ex (,,, : vii x.1955 1972). : 1ex ( : 10.viii.1973). : 5ex (, : vi viii.1964 1970). : 17ex (,,,,,, : vii viii.1967 1978). : 707, 6.9. 10.5. (vi- 30, vii- 498, viii- 157, ix- 20, x- 2ex). : Kamijo, 1936: 86; Nakayama and Okamoto, 1940: 197; Gu, 1973: 79. 41. Anomala japonica Arrow, 1913: 401 ( 4-38, 13-38, 18-38) Anomala japonica Arrow, Ann. Mag. Nat. Hist., (8)XII: 401, 1913: Kobayashi, 1985: 403; Kim, 1998a: 304

: : 105 ( ); 2001b: 115. : Kangwho (=? ). (A. mongolica), (A. albopilosa)..,..,. 2, 2... Ad:,., LPMB=1: 1.3: 1.2. BL: 20 26 mm, BW: 11.5 15 mm. :,,. :,,. : NK-. SK-GW,,,,,,, GS,,,,,,,,,, CN,,, GBN,,,, :,,,,,,,,,,,,, :. (130ex): [SWU] : 2ex ( : vi, viii.1984, 1985). : 18ex (,,,, : vi viii.1977 1990); 1ex ( : 3.vii.2007). : 2ex ( : viii.1987, 1993). : 1ex ( : 15.vii.1981). : 12ex (,,, : vi viii.1982 1997). : 1ex (8.viii.1989); 1ex ( : 31.vii.2002). [NAAS] : 2ex ( : 26.viii.1975). [SNU] : 1ex ( : 5.viii.1995). : 1ex ( : 24.viii.1988). : 1ex ( : 20.viii.1995). [KU] : 1ex ( : 10.viii.1974). : 3ex ( : viii.1928, 1971). : 1ex (17.vii.1990). [EWU] : 23ex (,,,, : vii viii.1970 1994). : 9ex (,,,, : vii ix.1955 1971). : 25ex (, : vii viii.1966 1971). : 22ex ( : vii.1968 1976). : 3ex (,, : vii.1960 1981). : 124, 6.12. 10.21. (vi- 7, vii- 90, viii- 25, x- 4ex). : Kolbe (1886) Ohaus (1915b) Euchlora cuprea Hope var. viridana Anomala mongolica. 42. ( ) Anomala mongolica (Faldermann), 1835: 379 ( 5-39, 13-39, 18-39) Euchlora mongolica Faldermann, Mem. Acad. St. Petersb., 2: 379, 1835.

106 Anomala mongolica: Bates, 1888: 374; Stebnicka, 1980: 209; Kim et al., 1991b: 68; 1991d: 193; 1991e: 179; 1992b: 154; 1994b: 111; 1996a: 59; 1996b: 128; 1997: 231; 2000: 132; 2002: 120; 2003: 126; 2004: 117; Kim, 1992: 106; 1993: 65; 1995a: 165; 1996c: 174: 1998a: 304; 2001b: 116; ESK and KSAE, 1994: 152. : (Kim and Park, 1991d; Kim, 2001b).,,,.,.,, (A. japonica). 2, 2. Ad:,,. LPMB=1: 1.1: 1.13. BL: 17 22 mm, BW: 9.7 12.5 mm. : Medvedev, 1952: 60; Zhang, 1984: 52; Liu et al., 1997: 38. :,,,. : NK-HB, YG, PB, PY,, WH,,, GW,?, SK- 80 120 1,000. :,,,,. (1,057ex): [SWU] : 37ex (,,,,,,,, -G,,,, : vii viii.1976 1999); 1ex ( : 20.vii.2009). : 76ex [,,,,,,,,,,, (8 ),,, : vi x.1920 1998]. : 19ex (,,,,,,,,,,, : v viii.1964 2003). : 81ex (,,,,,,,,, : vi viii.1978 1998). : 6ex (,,,,, : vi ix.1986 1995). : 2ex (vii.1990, 1991). [NAAS] NK-2ex ( : vii, viii.1921, 1922). : 12ex (,,,,, : vi ix.1987 1994). : 67ex (, : vi viii.1921 1996). : 17ex (,,, : vi viii.1974 1997). : 459ex (,, : vi ix.1988 1993); 1ex ( : vi.1991). : 27ex (,,,,, : vi viii.1922 1992). : 2ex (v, ix.1974, 1988). [SNU] : 1ex ( : 7.viii.1957); 3ex (vi, vii.1997). : 5ex (,, : vii ix.1964 1997). : 1ex ( : 28.viii.1997). : 2ex ( : vii.1997). : 18ex (,,,, : v viii.1976 1997). [KU] NK-2ex ( : 6.viii.1933); 1ex (, : 27.vii.1933). : 7ex (,,, : vii ix.1931 1984). :(,,, ). [EWU] : 5ex (,,, : vii viii.1973 1982). : 2ex (, : vi, vii.1984, 1996). : 1ex ( : 23.vi.1973). [EWU] : 3ex (, : vii, viii.1967, 1994). : 98ex (,,,,,,, : iv x.1954 1993). : 2ex (, : vii, viii. 1969, 1973). : 5ex (,,., : vii, viii.1961 1966, 1986). : 4ex (, : vii viii.1959 1967). [HSU] : 1ex ( : 23.vi.1997). [HMNH-NK] : 4ex ( : 13, 14.vii. 1982); 10ex (8 12.viii.1991). : 1ex ( : 10.vii.1982). : 4ex ( : 4 6.viii.1975); 23ex

: : 107 (9 11.vii.1977); 12ex (22, 23.vii.1982); 2ex (1.viii.1991). : 5ex (, : 31.vii.1982); 1ex ( : 29.viii.1989). : 2ex ( : 29.vii.1982). : 943, 5.12. 9.9. (v- 8, vi- 82, vii- 693, viii- 139, ix- 19, x- 1ex). : Liu et al., 1997: 38 ( ). : Mimela. A. viridana. 2 ( ),. 43. Anomala geniculata (Motschulsky), 1866: 171 Rhinoplia geniculata Motschulsky, Bull. Soc. Nat. Hist. Moscou, 39: 171, 1866. Anomala geniculata: Murayama, 1935: 5; KSPP, 1972: 206; Gu, 1973: 79 ( ); Stebnicka, 1980: 209; Kim, 1998a: 304, 2001b: 118. Anomala pleurimago Reitter, 1903; Best.-Tab., Brünn, 51: 72 (Korea), (syn. by Nomura, 1960): Niijima and Kinoshita, 1923: 97; Miwa and Chûjô, 1939: 38; Cho, 1957: 120; KZS, 1968: 135; Stebnicka, 1980: 209. : (KSPP, 1972); pleurimago= (KZS, 1968). : geniculata-. Pleurimago- Holotype, (Kiushiu); Paratype,.,.. 4,. BL: 11 16 mm. :,,. :?. : 1, Korea, 1893, Reitter (HMNH, A. pleurimago.). : Hungary, Reitter (Mimela costata). 44. Anomala chamaeleon Fairmaire, 1887: 317 ( 5-40, 13-40, 19-40) Anomala chamaeleon Fairmaire, Rev. d Ent., VI: 317, 1887: Heyden, 1889: 662; Niijima and Kinoshita,

108 1923: 114; 1927: 38; Maruta, 1929: 369; Haku, 1937: 121; Miwa and Chûjô, 1939: 34; Cho, 1957: 291; 1969: 676 ( ); KZS, 1968: 134; Kim and Kim, 1974a: 229; 1974b: 108; Kim and Nam, 1982a: 156; 1982b: 276; 1984a: 103; 1984b: 328; Kim, 1978: 349; Yoon and Nam, 1979: 149; 1986: 160; Yoon et al., 1990b: 111; Stebnicka, 1980: 209; Nam and Kim, 1983: 128; Kim et al., 1984: 169; 1991b: 68; 1991c: 154; 1991d: 193; 1991e: 179; 1994b: 111; 1996a: 59; 1996b: 128; 1997: 231; 1998: 170; 1999: 130; 2000: 132; 2001: 132; 2002: 120; 2003: 126; Park and Kim, 1993: 113; Kim, 1992: 106; 1993: 65; 1995b: 140; 1996c: 174: 1997: 231; 1998a: 305; 2001b: 118; ESK and KSAE, 1994: 152; Lie et al., 1997: 41. Anomala dubia Ballion, Bull. Moscou., IV: 344, 1870: Bates, 1888: 374; Kim, 1998a: 305 (syn. nov.). Anomala geniculata (Motschulsky, 1866) ( ): Kim and Nam, 1981: 125. :.,.,.,, 2/3.,. 3, 3. Ad:,,., LPMB=1: 0.6: 0.9. BL: 12 17 mm, BW: 7 13 mm. : Murayama, 1931: 59; Medvedev, 1952: 63. :,,. :,,,. : NK-HB,, YG,,,, HN,,,, PB,,,, PY,,,,, PN, WB, WN, GW,, GG,? Soktan, Tschonbonsan, Kuson, Nyompyang, Kekki, SK- (5 ) 120 160 5,000. (3,941ex): [SWU] : 175ex (,,,,,,,,,,,,,, : vi viii.1958 2001); 2ex ( / : 10/20.vii.2009). : 403ex (,,,, 12 : iv x.1953 1999); 1ex ( : 28.vii.2000); 3ex ( : 23.vii.2005). : 135ex (,,,,,,,,,,,,,,,,,,, : v ix.1944 2002). : 278ex (,,,,,,,,,,,,,,, : iv ix.1980 2000). : 46ex (,,,,,,,,,, : vi viii.1983 1998). : 13ex (viii. 1955 1999). [NAAS] NK-6ex (,,, : vii x.1924 1931). : 13ex (,,,,, : vi viii.1923 1994). : 551ex (,,,,,,, : v ix.1921 1996). : 96ex (,,,,,,,,, : vi viii. 1975 1996). : 782ex (,,, : vi ix.1980 1993); 2ex ( : vii.1991). : 98ex (,,,,,,, : vi viii.1922 1994). : 10ex (v viii.1919 1996).

: : 109 [SNU] : 11ex (,, : vii, viii.1958 1997). : 68ex (,,,,,,,,,,, : v x.1964 1997). : 5ex ( : vii.1983); 1ex ( : 29.vi.1994). : 3ex (,, : vii, viii.1995 1997). : 169ex (,, : v ix.1968 1997). : 12ex (vii.1971).?: 4ex ( : 5.viii.1993). [KU] NK-5ex ( : 16.vii.1930); 2ex ( : 4.viii. 1933). : 101ex (Mt.,,,,,,, : vi viii.1922 1994). : 90ex [,,, (6 ),,,, : iv ix.1933 1993]. : 164ex (,,,, : 1957 1988). : 107ex (,,,,,,, : iv viii.1960 1990). : 18ex (,,, : vii viii.1974 1979). : 8ex (vii viii.1955 1990). [EWU] : 30ex (,,,,,, : vii ix.1960 1994). : 116ex (,,,,,,,, : iv x.1954 1998). : 12ex (,,,,, : vii viii.1960 1973). : 13ex (,,, : vii viii.1964 1975). : 11ex (,,,,,,,, : vii viii.1959 1978). : 11ex (v viii.1957 1987). [HSU] : 22ex ( : vi.1994 1997). [HMNH-NK] : 1ex ( : 22.vii.1975). : 4ex ( : 13 14.vii.1982); 8ex (8 12.viii.1991). : 1ex (16.vii.1975); 3ex (, : 8, 10.vii.1982). : 4ex ( : 30.vii.1974); 47ex (4 8.viii.1975); 15ex (9 11.vii.1977); 9ex (22 25.vii.1982); 19ex (1 4.viii.1991). : 2ex ( : 31.vii.1982); 1ex ( : 29.viii.1989). : 1ex ( : 29.vii.1982). : 1,975, 5.2. 10.23. (v- 5, vi- 94, vii- 1,106, viii- 675, ix- 83, x- 12ex). : Murayama, 1936: 1 6; 1938: 259 264; Gu, 1973: 78. :, 1. 45. Anomala rufocuprea Motschulsky, 1860: 14 ( 5-41, 13-41, 19-41) Anomala rufocuprea Motschulsky, Etud. Ent., 9: 14, 1860: Heyden, 1887: 252; Bates, 1888: 373; Masaki, 1936: 260; Kim, 1956: 9; 1958: 98; 1960: 27; Cho et al., 1967: 198; Hyun and Woo, 1970: 80; Gu, 1973: 78; Kim et al., 1976: 102; Stebnicka, 1980: 209; Park and Park, 1987: 673; Yoon et al., 1990a: 140 ( ); Park et al., 1993: 178; Lee et al., 1985: 418; Kim, 1998a: 305; 2001b: 120; ESK and KSAE, 1994: 152. Anomala motschulskyi Harold, Deut. Ent. Zeit., 21(2): 351, 1877 (Japan): Niijima and Kinoshita, 1923: 108; Okamoto, 1924: 176; Yoshino, 1935a; 14; Morita, 1936: 861; Miwa and Chûjô, 1939: 37; Kim, 1958: 91 ( ); Cho, 1957: 291; 1963: 219; Cho et al., 1968: 264; Hyun and Woo, 1969: 192; Won and Choi, 1968: 379; Nam and Kim, 1983: 128. Anomala motschulskyi viridicuprea Ohaus, Stett. Ent. Zeit., 76: 321, 1915 (Japan): Niijima and Kinoshita, 1923: 108; Miwa and Chûjô, 1939: 37; Cho, 1957: 291. Anomala chamaeleon Fairmaire, 1877 ( ): Kim and Kim, 1976: 102; Kim et al., 1979: 84; 1987b: 224;

110 1991b: 68; Nam and Kim, 1983: 128; Yoon et al., 1990a: 140. : rufocuprea= (Kim, 1960 2), (Hyun and Woo, 1970), (Gu, 1973). :,.,..., 2/3. 3 3. Ad:,., LPMB 1: 0.95: 1.05. BL: 12 16 mm, BL: 7 10 mm. : Tamari et al., 1985: 359 361; Nakano et al., 1986: 254 259. :,,. :,. : NK-, SK-GW-,,, GS,,,, GN,,,, JN, ( A. chamaeleon, ). (361ex): [SWU] : 1ex ( : 4.viii.1998). : 3ex (, : vi, vii.1936, 1998). : 1ex ( : 31.vi.1998). : 220ex ( : 15.viii.1988, Yeo S.D.); 130ex ( : 10.viii.1988, Yeo S.D.); 1ex ( : 10.viii.1988). [SNU] : 1ex ( : 21.vi.1992); 1ex ( : 3.ix.1988). : 2ex ( : viii.1988, 1994). [EWU] : 1ex ( : 17.viii.1945(?)). : Gu, 1973: 78; Kim and Kang, 1993: 415; Setokuchi and Ouchi, 1982: 273 (Kyushu). 46. ( ) Anomala koreana Kim, 1997: 279 ( 5-42, 13-42, 19-42) Anomala koreana Kim, Kor. J. Entom., 27(4): 279, 1997 ( ): 1998a: 305, 2001b: 121..,,,,,...,.,,.,.,

: : 111... Ad:.,. LPMB=1: 0.9: 1.3. BL: 10 13 mm. BW: 5.8 7 mm. :. :. :,. (18 7ex): [SWU] : 1 [ : 20.vii.1993, E.H. Kim ( )]; 1 3 [ : 20.vii.1993, E.H. Kim ( )]. : 1 ( : 15.viii.1993, J.Y. Lim); 1 ( : 13.vii.1987, S.Y. Ku). : 1 1 ( : 9.vii.1987, K.H. Hong); 1 ( : 2.vii.1981, K.H. Oh); 1 ( : 9.x.1984, H.S. Kim). : 1 2 ( : 24, 25.vii.1931, P.S. Cho); 1 1 ( : 13.vii.1985, E.Y. Lee). : 2 ( : 24.vii.1956, S.W. Park)-. [KU] : 1ex ( : 18.vii.1976). [EWU] : 1ex ( : 16.vii.1975); 4ex ( : vi viii. 1960 1971). : 1ex ( : 24.vii.1971). 47. Anomala luculenta Erichson, 1847: 620 ( 5-43, 13-43, 19-43) Anomala luculenta Erichson, Naturg. Ins. Deut., III: 620, 1847; Ohaus, 1915b: 316; Niijima and Kinoshita, 1923: 99; Maruta, 1929: 369; Morita, 1936: 860; Eguchi, 1932: 54; Kamijo, 1932: 22; 1936: 86; Mochizuki, 1935: 32; 1936: 211; Mochizuki and Tsunekawa, 1937: 90; Haku, 1937: 121; Masaki, 1936: 260; Miwa and Chûjô, 1939: 37; Mori and Cho, 1940: 13; Cho, 1934: 75; 1947: 65; 1957: 219; 1969: 678; KZS, 1968: 135 ( ); Kim, 1978: 349; Stebnicka, 1980: 277 ( ); Kim et al., 1991b: 68; 1996a: 59; 1999: 130; 2000: 132; 2002: 120; 2003: 126; Kim, 1992: 106; 1993: 65; 1998a: 305; 2001b: 121; ESK and KSAE, 1994: 152; Lie et al., 1997: 41. Anomala luculenta purpurea Niijima and Kinoshita, Res. Bull. Exp. Forest, Hokkido Univ., II (2): 99, 1923 (Corea): Haku, 1937: 121; Miwa and Chûjô, 1939: 37; Cho, 1957: 219. Anomala luculenta smaragdina Ohaus, Stett. Ent. Zeit., 76: 317, 1915 ( ). Anomala daurica rufofemorata Reitter, Ver. Nat. Ver. Brünn, Best-Tab., 51: 74, 1903 ( ). Anomala lucens Ballion, 1871 ( ): Kim, 1981: 344; Kim et al., 1987c: 506; 1991e: 179; 1992b: 154; Yoon et al., 1990b: 111. Anomala puncticollis Harold, 1877 ( ): Kim and Chang, 1987a: 104. : (Cho, 1947); lucens= (Yoon et al., 1990b).,,,

112,..,.,,.,..,. Ad: 1, W. LPMB=1:1.1:1.2. BL: 13 17 mm, BW: 7.5 9.5 mm. : Zhang, 1984: 52. :,,,. :,,,,. : NK-HB,, HN,, PB,,, PN,,,, WH, GW,, GG,? Nyompyang, Kekki (Vekki), Kuson, Chon-Ge-Charzon Wald, SK-GW,,,,,,,,,,,, GS,,,,,,,,,,,,,,,,,, (12 ),,,,,,,,, CBN,,,,,,,,, GBN,,,,,,,,,, JBN,,,,,,,,?,, Koshou, Shoko. (683ex): [SWU] : 50ex (,,,,,, : v ix.1962 2001). : 82ex [,,,,,,, (8 ),,,, : iv ix.1928 1998]. : 113ex (,,,,, : vi viii.1960 1996); 1ex ( : 27.vii.2008). : 15ex (,,,,, : vi viii.1979 1998); 2ex ( : 4.viii.2001). : 3ex (, : vii viii.1983, 1997). : 1ex (viii.1983). [NAAS] NK-1ex (?: 10.viii.1921). : 3ex (,, : vi, vii.1980 1990). : 7ex (, : vi viii.1922 1989). : 1ex ( : 23.vii.1987). : 1ex ( : 10.vii.1993). : 2ex ( : vii.1975). [SNU] : 5ex (,,, : vii, viii.1974, 1977, 1997). : 53ex (,,,,,,, : iv ix.1968 1995). : 2ex (, : vi, viii.1983, 1988). : 1ex ( : 24.vii.1990); 9ex ( : vii viii.1973 1987). [KU] NK-1ex ( : 14.vii.1932). : 30ex (,,,, : vi, vii.1927 1981). : 98ex [,,,,,, (3 ),, : v ix.1933 1997]. : 16ex (,,, : vii viii.1957 1982). : 54ex (,,,, : vi viii.1960 1990). : 3ex (, : viii.1970, 1974); 11ex ( : vi viii.1961 1986). : 1ex (30.vii.1955).?: 1ex (Koshou: 15.viii.1936); 1ex (Shoko: 6.viii.1916). [EWU] : 5ex (, : vi viii.1973 1983). : 104ex (,,,,, : iv x.1954 1984). : 2ex (, : vii, viii.1961, 1967). : 3ex ( : viii.1975). [HMNH-NK] : 1ex ( : 5.vii.1991). : 1ex ( : 5.vii.1977). : 573, 4.18. 9.28. (iv- 2, v- 17, vi- 173, vii- 210, viii- 153, ix- 20ex). : Eguchi, 1932: 54; Kamijo, 1936: 86; Liu et al., 1997: 41.

: : 113 : Stebnicka (1980) (A. ogloblini). 48. Anomala corpulenta Motschulsky, 1853: 28 ( 5-44, 13-44, 19-44) Anomala corpulenta Motschulsky, Etud. Ent., II: 28, 47, 1853: Niijima and Kinoshita, 1923: 95; Okamoto, 1924: 176; Maruta, 1929: 369; Kamijo, 1932: 21; Miwa and Chûjô, 1939: 34; Cho, 1957: 291; 1969: 677 ( ); Kim and Kim, 1975: 244; 1977: 131; Kim and Nam, 1982a: 156; 1982b: 276; Kim, 1978: 351; Yoon and Nam, 1979: 149; 1986: 160; Yoon et al., 1990b: 111; Stebnicka, 1980: 277; Park and Park, 1986: 128; 1992: 178; Kim et al., 1991b: 68; 1991e: 179, 1994b: 111; 1996a: 59; 1997: 231; 1998: 170; 2001: 132; 2002: 120; Kim, 1992: 106; 1993: 65; 1995a: 165; 1997: 231; 1998a: 306; 2001b: 123; ESK and KSAE, 1994: 152; Lie et al., 1997: 40. Anomala gottschei Kolbe, Arch. f. naturf., 52: 190, 1886 ( ): Reitter, 1903: 71; Okamoto, 1924: 176. Anomala geniculata Motschulsky, Bull. Soc. Hist. Nat. Moscou, 39: 171, 1866 (Japan): Lewis, 1896: 334. Anomala daimiana Harold, 1877 (?): Okamoto, 1924: 176; Eguchi, 1932: 56; Murayama, 1937a: 35; Miwa and Chûjô, 1939: 34; Cho, 1957: 119, 1963: 219; Ku, 1963: 28; 1973: 77; Won and Choi, 1968: 379; KZS, 1968: 135; KSPP, 1972: 206; Yoon and Nam, 1978: 83; 1986: 160; Kim and Nam, 1978: 134; Kim et al., 1987c: 506; 1991b: 68; 1991d: 193; ESK and KSAE, 1994: 152. : daimiana= ( ) (Cho, 1963 9). :,.,,.,..,.,.. Ad:,. LPMB=1: 1.2: 1.4. BL: 14 18 mm, BW: 8 10 mm. : Murayama, 1931: 52; Medvedev, 1952: 62; Zhang, 1984: 54. :,,. :,,,. : NK-HN, PB, PY,, WN, GW,,? Kuson, Tschonbonsan, SK- ( 30 ) 110 140 1,500.

114 (1,033ex): [SWU] : 12ex (,,,, : v ix.1981 1997). : 134ex [,,,,,,,,,,, (16 ),,, : v x. 1934 1998]. : 2ex ( : 18.vi.2008). : 59ex (,,,,,,,,,, : v x.1981 1998). : 17ex (,,,,,, : vi ix.1986 1992); 2ex ( : 5.vi.2008). : 6ex (,, : vii ix.1981 1995). [NAAS] : 5ex (,, : vi viii.1983 1992). : 314ex ( : v ix.1921 1997); 5ex (,, : vi ix.1986 1996). : 29ex (,, : vi ix.1975 1991). : 17ex (,,,, : vi, vii.1991 1993). : 8ex (,,, : vii ix.1975 1996). : 6ex (v viii.1919 1988). [SNU] : 2ex (, : ix.1993, viii.1996). : 203ex [,,,,,,, (3 ),,,,, : iv x.1966 1997]. : 5ex (,,, : vi vii.1983 1995). : 16ex (,, : vi viii.1978 1995). [KU] : 3ex ( : 6.vii. 1929); 1ex ( : 21.vi.1984). : 53ex [,,,,, (7 ),, : iv ix.1933 1988]. : 3ex (,, : vii.1957,1979). : 2ex ( : 11.vii.1977). : 6ex (, : v vii.1983 1985). [EWU] : 2ex (, : v, vi.1973, 1977). : 102ex (,,,, : iii x.1960 1994). : 4ex (,,, : vi viii.1967 1982). : 1ex ( : 4.viii.1965). [HSU] : 3ex ( : vi.1994 1997). [HMNH-NK] : 1ex ( : 8.viii. 1991). : 4ex ( : 16.vii.1975); 3ex (29.vi.1991); 2ex ( : 8.vii.1982); 1ex ( : 10.vii.1977). : 857, 4.25. 10.24. (iv- 2, v- 24, vi- 277, vii- 380, viii- 138, ix- 25, x- 11ex). : Murayama, 1938: 259; Nakayama and Okamoto, 1940: 197; Kim and Kang, 1993: 415; Liu et al., 1997: 40. 49. Anomala ignicolor Reitter, 1903: 73 Anomala ignicolor Reitter, Verh. Nat. Ver. Brünn, Best.-Tab., 51: 73, 1903: Niijima and Kinoshita, 1923: 114; Miwa and Chûjô, 1939: 36; Cho, 1957: 119; KZS, 1968: 135 ( ); ESK and KSAE, 1994: 152; Kim, 1998a: 306; 2001b: 125. :. 13 mm,,.,..,,...,, 2/3.. 1 BL: 14.3 mm, BW: 8 mm.

: : 115 :. :?. : Korea, 1, 1898, Reitter. : Hungary, Reitter (Mimela costata Anomala geniculata ). 50. Anomala ogloblini S. Medvedev, 1949: 171 Anomala ogloblini S. Medvedev, Fn. URSS Col., 10(3): 171, 1949: Kalinina, 1989: 415; ESK and KSAE, 1994: 152; Kim, 1998a: 306 ( ); 2001b: 123. : (ESK and KSAE, 1994). (A. luculenta),,. Ad:. :,,. :?. :. : Stebnicka (1980) A. luculenta ( ) Kalinina (1989) ogloblini. Stebnicka.. 51. Anomala cuprea Hope, 1839: 72 Euchlora cuprea Hope, Proc. Zool. Soc., VII: 72, 1839. Anomala cuprea: Miwa and Chûjô, 1939: 34; Kim, 1956: 9; 1958: 98; Cho, 1957: 119; 1969: 677 ( ); Ku (Gu), 1963: 28; 1973: 77; Kim, 1978: 354; Stebnicka, 1980: 276; Kim and Kim, 1982b: 276; 1982a: 156; Kim and Park, 1991d: 193; ESK and KSAE, 1994: 152; Kim, 1998a: 304; 2001b: 117. Anomala (Euchloomala) cuprea: Stebnicka, 1980: 276. Euchlora cuprea Hope var. viridana Kolbe, Arch. f. naturf., 52: 192, 1886 ( ).

116 : (Gu, 1973). :. (A. mongolica),.. Ad:,. LPMB=1:1.1:1.0. BL: 17 25 mm. :,,,. :,. : NK-HN,, PN,, PY, GW, SK- GW, GS,,,. :. :, (A. mongolica). Anomala (= ) 1. Anomala corrugata Bates, 1866 Anomala corrugata Bates, Proc. Zool Soc, Lond.: 343, 1866; Niijima and Kinoshita, 1923: 114. 2. Anomala multistriata Motschulsky, 1861 Hetroplia multistriata Motschulsky, Etud. Ent., X: 7, 1861: Nakane, 1972: 427 Tsushima Korea. 3. Anomala polyanor Ohaus, 1916 Anomala polyanor Ohaus, Arch, f, Nat., 83, A4: 4, 1916: Niijima et Kinoshita, 1927: 132; Miwa and Chûjô, 1939: 38; Cho, 1957: 120; KZS, 1968: 135 ( ) Blitopertha.

: : 117 4. Anomala schoenfeldti Ohaus, 1915 Anomala schoenfeldti Ohaus, Stett. Ent. Zeit., 76: 332, 1915 ( ): Murayama, 1935: 5; Miwa et Chûjô, 1939: 39; Cho, 1957: 120; KZS, 1968: 135 ( ), Kim and Lee, 1991b: 68; ESK and KSAE, 1994: 152 Murayama. 5. Anomala spiloptera Burmeister, 1855 Anomala (Idiocnema) spiloptera Burmeister, Handb. IV. 2: 500, 1855: Stebnicka, 1980: 209 ( ). 6. Anomala viridana Kolbe, 1886 Euchlora viridana Kolbe, Arch. f. Naturg., LII: 192, 1886 (Korea): Cho, 1947: 65 ( ); 1963: 62 ( ); 1969: 678 ( ); KSPP, 1972: 206 ( ) 39, ( 30 ) 100 A. mongolica A. japonica. Genus Bifurcanomala Kim, 1998b: 312. 10, (Kim, 1998b) Anomala.. : Melolontha aulax Wiedemann, 1823. Zool. Mag., II(1): 93. : ( 1 ). :. : Kim (1998b).

118 52. Bifurcanomala aulax (Wiedemann), 1923: 93 ( 5-45, 13-45, 19-45) Melolontha aulax Wiedemann, Zool. Mag., II: 93, 1923. Anomala aulax: Stebnicka, 1980: 209; Kim, 1993: 65; 1995b: 140; 1997: 231; 1998c; 357; Kim and Park, 1991d: 192; Kim and Lee, 1997a: 231; Lee et al., 1994: 147; ESK and KSAE, 1994: 152. Bifurcanomala aulax: Kim, 1998b: 312 ( ); 2000: 132; 2001b: 126. Anomala costifera Reitter, Wien. Ent. Zeit., 14: 209, 1895 (Corea): 1903: 66; Niijima and Kinoshita, 1923: 92; Okamoto, 1924: 175; Murayama, 1937b: 14; Miwa and Chûjô, 1939: 34; Cho, 1957: 291; KZS, 1968: 134; KSPP, 1972: 205; Gu, 1973: 77; Stebnicka, 1980: 209; Kim and Lee, 1991b: 68; Kim, 1992: 106; ESK and KSAE, 1994: 152. : (ESK and KSAE, 1994); costifera= (KZS, 1968), (KSPP, 1972)..,...., 10,. 2 4. Ad:.,... BL: 11 16 mm, BW: 5.8 9 mm. : Zhang, 1984: 49 (A. aulax). :,,. :,,,. : NK-? Soktan, Tschon-bon-san, SK- (15 ) 60 80 800. (781ex): [SWU] : 7ex (,,,, : vi viii.1934, 1984 1997). : 117ex [,,,,,,,, (6 ),, : vi x.1972 1998]; 1ex ( : 8.vi.2003). : 6ex (,,,,, : v vii.1987 1999). : 19ex (,,,,,, : vi vii.1976 2000). : 77ex (,,,,,,, : vi vii.1984 1998). : 32ex (vi vii.1983 1994); 1ex (10.vii.2002). [NAAS] : 2ex ( : vi.1967, 1984). : 1ex ( : 25.vii.1983). : 2ex (, : vi, vii.1973, 1975). : 30ex (v viii.1974 1992). [SNU] : 30ex [,, (4 ),,,, : vi ix. 1955 1992]. : 3ex ( : vi x.1983 1988). : 6ex ( : vi vii.1994 1997). : 61ex (

: : 119,, : vi viii.1926 1995). : 1ex (14.vii.1965). [KU] : 3ex ( : vi.1979, 1984). : 137ex [,,,,,,,,, (6 ): iv ix.1959 1996]. : 1ex ( : 28.vii.1979). : 13ex (,, : vi vii.1977 1990); 124ex ( : vi.1973). : 52ex (,,,, : vi vii.1974 1993). : 3ex (vii.1962 1972). [EWU] : 8ex (,,, : v vii.1954 1985). : 1ex ( : 20.vii.1969). : 1ex ( : 16.vi.1979). : 30ex (,,, : vi vii.1978 1992). : 4ex (vii.1965 1987). [HSU] : 6ex ( : v vi.1994 1997). : 640, 4.20. 11.23. (iv- 3, v- 13, vi- 430, vii- 183, viii- 3, ix- 2, xi- 2ex). : Nakayama and Okamoto, 1940: 198 (=costifera); Gu, 1973: 77. Genus Chejuanomala Kim, 1998b: 313.,.. (1998b) Anomala, 1. : Anomala quelparta Okamoto, 1924. Bull. Agr. Exp. Sta., Gov. Gen. Chosen, 1(2): 177. : ( 1 ). :. : Kim (1998b). 53. Chejuanomala quelparta (Okamoto), 1924: 177 ( 5-46, 13-46, 19-46) Anomala quelparta Okamoto, Bull. Agr. Exp. Sta., Gov. Gen. Chosen, 1(2): 177, 1924: Niijima and Kinoshita, 1927: 35; Miwa and Chûjô, 1939: 39; Cho, 1957: 120; 1963: 219 ( ); 1969: 679; Cho et al., 1968: 265; Kim, 1978: 349; Lee et al., 1985: 418; ESK and KSAE, 1994: 152. Chejuanomala quelparta: Kim, 1998b: 313; 2001b: 128. : Saishuto (= ). :.,.,,.

120.,.,.. 2 4.. Ad:,. BL: 14 16 mm, BW: 7 8.5 mm. :. :. :. : JJ. (138 ): [SWU] : 1ex (16.vi.1963); 6ex ( : 14.vii.1997); 2ex ( : 14.vi.1994). [NAAS] : 25ex (v vi.1987 1996). [SNU] : 5ex ( : 1994). [JM] : 1ex ( : 22.vi.1981); 8ex (20, 29.vi.1983); 56ex (10 25.vi.1993); 2ex (8, 14.vii.1993); 4ex ( : 3.vi.1992, 27.vi. 1993); 2ex ( : 3.vii.1982); 2ex ( : 4.vi.1983); 1ex ( : 9.vi.1994); 22ex (,,,,, : vi vii.1993). [EWU] : 1ex ( : 9.vii.1965). :. Family Dynastidae MacLeay, 1819: 22. Dynastinae MacLeay, 1819. (Melolonthidae),,..,. (Ocular canthus) Cryptodus. 3 8 10.,. 1,. Endrödi (1985) 7 1,366 6 651, 3 29. Gottsche (1884) Kolbe (1886) Phileurus chinensis. Heyden (1887) P. morio. Niijima and Kinoshita (1923) Oryctes rhinoceros, (Kim, 2001b). Kôno (1931) (Trypoxylus Allomyrina dichotoma). 60 1990 Pentodon quadridens.

: : 121 (Kim, 1997),. 3, 50. : Dynastes MacLeay, 1819: 22. : 8 (Endrödi, 1985) ( 3 ). : 103 (Endrödi, 1985) ( 3 ). : 2,000 1,366 (Endrödi, 1985) ( 3 ). :. : Reitter (1898), Nijiima and Kinoshita (1923, 1927), Endrödi (1985), Kim (1997b, 2001b); Phileurini=Endrödi (1977, 1985); Trypoxylus=Kôno (1931), Endrödi (1985). 1., 35 55 mm.,., Allomyrina dichotoma, 25 mm 2 2., 2 Pentodon quadridens, 1,., Eophileurus chinensis ( ) Tribe Dynasyini MacLeay, 1819: 217.,,.. 10. : Dynastes MacLeay, 1819. : 10 (Endrödi, 1985) ( 1 ). : 60 (Endrödi, 1985) ( 1 ). :,. : Endrödi (1985).

122 Genus Allomyrina Arrow, 1911: 153. : Scarabaeus dichotoma Linné, 1771. : 3 (Endrödi, 1985) ( 1). : (2 ), (1 ). : Endrödi (1985). 54. Allomyrina dichotoma (Linné), 1771: 527 ( 5-47, 13-47, 19-47) Scarabaeus dichotoma Linné, Mant. II: 527, 1771. Trypoxylus dichotoma: Kôno, 1931: 159; Kamijo, 1932: 20. Xylotrupes dichotomus: Cho, 1933: 81; 1936: 25; 1956: 76; 1963: 221; 1969: 686; Mochizuki, 1936: 210; Morita, 1936: 860; Mochizuki and Tsunekawa, 1937: 92; Mori and Cho, 1940: 13; Cho et al., 1967: 198; 1968: 265; KZS, 1968: 138; Hyun and Woo, 1970: 80; Kim and Kim, 1971: 160; 1975: 252; Kim and Nam, 1983: 87; Yoon et al., 1990: 128. Allomyrina dichotoma: Miwa and Chûjô, 1939: 75; Cho, 1947: 65 ( ); 1957: 301; Kim and Kim, 1974b: 108; Kim and Nam, 1982a: 276; 1982b: 156; 1984: 328; Kim, 1978: 372; Stebnicka, 1980: 210; Yoon and Kim, 1981: 47; Nam and Kim, 1983: 128; Endrödi, 1985: 627; Yoon et al., 1990b: 111; Kim, 1992: 106; 1993: 65; 2001b: 80; Park et al., 1993: 177; Park and Kim, 1993: 113; ESK and KSAE, 1994: 152. :, 2. 2, 2. 10., 2,... :.,.. 2 (, spur= ). BL: 35 53 mm, BW: 18 27 mm, : 9.5 32 mm, : 5 15 mm.

: : 123 :,,,,. :,,. : NK-HB,, HN, PB,, PY, WH,,,,,,, GW, SK-,,,,,,,,, CB,,, GB,,,,,, GN,,,, JB,,,,,,, JN,, JJ. (51ex): [SWU] NK, : 1 1 ( : 3.vii.1991). : 1ex ( : 31.vii.1980). : 1 ( : 1.viii.1999). : 1 ( : 31.vii.1981); : 2ex (, : 4.viii.1980, 1992). : 6ex (v viii.1974 1999). [NAAS] : 1 ( : 27.ix.1993). : 10ex (, : vii ix.1993). : 1 ( : 12.vii.1991). : 2 ( : 9, 23.vii.1996). [SNU] : 8ex ( : vii, viii.1973 1979). : 2ex (29.vii.1971, 30.vii.1993). [EWU] : 1ex ( : 19.v.1954). : 2ex ( : 23.vii. 1986); 2ex ( : 29.vii.1970). : 1ex ( : 4.viii.1965). : 4ex (vi, vii.1964 1973). [HMNH- NK] : 2ex ( : 5 7.vii.1991); 2ex (8 12.viii.1991). :,. ( ) Tribe Phileurini Burmeister, 1847: 138. Phileuridae Burmeister, 1847. 1 2. (mentum),. 10 9 8.,. 3 4.. : Phileurus Latreille, 1807. : 35 (Endrödi, 1985) ( 1 ). : 183 (Endrödi, 1985) ( 1 ). :. : Endrödi (1985), Kim (2001b).

124 Genus Eophileurus Arrow 1908: 332., 1 2. 10,,. : Geotrupes platanus Wiedemann, 1823. Zool. Mag., II(I): 5. : 27 (Endrödi, 1985) ( 1 ). :. 55. Eophileurus chinensis (Faldermann), 1835: 370 ( 5-48, 6-48, 13-48, 19-48) Phileurus chinensis Faldermann, Mem. Acad. Petr., II: 370, 1835: Kolbe, 1886: 194; Maruta, 1929: 370; Morita, 1936: 860; Haku, 1936: 121; Mochizuki and Tsunekawa, 1937: 92; Kim and Kim, 1975: 252; Kim and Nam, 1982b: 276. Eophileurus chinensis: Miwa and Chûjô, 1939: 74; Cho, 1957: 301; 1963: 221 ( ); 1969: 685; Cho et al., 1968: 265; KZS, 1968: 138; Endrödi, 1977: 99, 1985: 663; Kim, 1978: 371; Yoon and Nam, 1979: 150; Stebnicka, 1980: 210; Kim and Nam, 1982b: 156; Kim et al., 1982a: 145; 1991b: 67; 1991e: 179; 1977: 234; 2001: 132; 2005: 79; Park et al., 1993: 178; ESK and KSAE, 1994: 152; Lie et al., 1997: 26; Kim, 1998c: 357; 2001b: 81. Phileurus morio Faldermann, Mem. Acad. Petr., II: 371, 1835: Heyden, 1887: 254; Okamoto, 1924: 181; Miwa and Chûjô, 1939: 74.,. 10,. :., 1.,... :,. BL: 18 24 mm, BW: 9 12 mm. :,,,. :,,,. : NK-HN, SK-GS,,,, (,,,,

: : 125,,, ),,,,,,,,,, CBN,,,, GBN,,,, JBN,,,,,,, JJ,. (72ex): [SWU] : 1ex ( : 2.viii.1982); 1ex ( : 5.v.1998); 2ex ( : 11.iv.1998). : 1ex ( : 10.viii.1987). : 2ex ( : 16.viii.2009); 2ex (10.x.2009). [NAAS] : 9ex ( : v viii.1926 1934, 1958 1992); 1ex ( : 19.viii.1976); 1ex ( : 15.vi.1993). : 4ex ( : vii, viii.1996); 1ex ( : 27.vi.1973). : 2 ( : 16.viii.2009); 1 1 (10.x.2009). : 1ex ( : 15.vii.1993). [SNU] : 13ex (,,, : v xi.1964 1992); 1ex ( : 25.v.1967). : 1ex ( : 22.vi.1992). : 1ex ( : 21.vi.1965); 1ex ( : 20.vii.1969); 1ex ( : 25.vi.1994). [EWU] : 1ex ( : 14.vii.1962); 2ex ( : vi.1957, 1962); 1ex (x.1966); 2ex ( : v.1962; vi.1961). : 1ex ( : 28.vii.1964). [KU] : 4ex ( : 18.vii. 1939); 2ex (1934, 1938); 1ex ( : 20.v.1989); 1ex ( : 16.x.1971); 1ex ( : 20.ix.1975). : 1ex ( : 29.vii.1957). : 1ex ( : 28.vii.1979). : 1ex ( : 20.viii.1956). : 1ex ( : 10.vii.1964). [HSU] : 2ex ( : 26.v., 8.vi.1997). ( ) Tribe Pentodontini Mulsant, 1842: 381. Pentodonaires: Mulsant, 1842: 381.,. 1 3.. : Pentodon Hope, 1837. : 22 (Endrödi, 1985) ( 1 ). : 500 (Endrödi, 1985) ( 1 ). :. : Endrödi (1985), Kim (1997b, 2001b). Genus Pentodon Hope 1837: 30. 7, 2 9, 6,

126, (Endrödi, 1985). P. quadridens 4,, distantides, mongolicum, bidentulum (Endrödi, 1985). bidentulum. : Scarabaeus punctatus Villers, 1789. Col. Man., Lond., 13: 92. 56. Pentodon quadridens (Gebler), 1845: 100 ( 6-49, 13-49, 19-49) Scarabaeus quadridens Gebler, Bull. Acad. St. Petersb., 3: 100, 1845. Pentodon quadridens: Kim, 1997b: 225 ( ); 2001b: 82; Kim and Lee, 1997a: 234.,.,,,. 10,. 2. 1 2, 3,.,,.. 3... BL: 20 mm, BW: 11.5 mm. :,,,,,,,. :. : SK-GG, CN,. (6ex): [SWU] : 2 ( : 23.vi.1996). [NAAS] : 4ex ( : vii, viii.1996). (= ) 1. Oryctes rhinoceros Linné, 1758 Oryctes rhinoceros Linné, Syst. Nat. ed. I: 346, 1758: Niijima and Kinoshita, 1923: 205; Miwa and Chûjô,

: : 127 1939: 74; Cho, 1957: 301; 1968: 138 ( ); Nomura, 1960: 73; KSPP, 1972: 208; Stebnicka, 1980: 210; Endrödi, 1985: 520. Family Melolonthidae Leach in Samouelle, 1819: 189.,,,..., 3. ( ) 6. (, spurs) 1 2,,.,. : Melolontha Fabricius, 1775: 31. : Gottsche 1884 ( ) 4 Kolbe (1886) 1 (Apogonia cupreoviridis), Serica (=Maladera) 2 Lachnosterna (=Holotrichia) 1. Heyden (1887) 8 6, 1910 Brenske, Reitter, Arrow. Niijima Kinoshita (1923, 1927) 23. 1930 Murayama (1954) 61. 90. (Kim, 2000) Holotrichia 15 10. Sericinae (Kim et al., 1991f, 1997b, 2002a, b, c) 45 19. (Kim, 2001b) 19 55, 19 53 4 57.. : 5 ( 3 ). : 7 28 ( 3 ). : 750 (Houston and Weir, 1992) ( 19 ). : 11,000 (Houston and Weir, 1992) ( 53+4 ). :. : Kolbe (1886), Heyden (1887, 1889), Bates (1888), Reitter (1902), Moser (1915a, 1919, 1920), Nijiima and Kinoshita (1923, 1927), Murayama (1954), Nomura (1973), Stebnicka (1980), Lie et al. (1997), Sabatinelli and Pontuale (1998), Kim et al. (1991f, 1997b, 2002a, b, c), Kim (2000, 2001b),

128 Smetana and Andrew (2006). : Murayama (1931), Medvedev (1952), Zhang (1984). 1. (, spur) 1. 2 1. Hopliinae 2 ( ), 2 2 2.,. Melolonthinae.. Sericinae Subfamily Hopliinae Latreille, 1829: 563. Hoplides Latreille, 1829.,. 9 10 3, (insertion).,. (, spur), 1. 1 2,. 2 10 2 3,. : Hoplia Illiger, 1803: 226. : 4 ( 2 ). : 250 (Hoplia 170 )( 3/4 ). :,. : Hardy (1977). 1., (gen. Ectinohoplia), 1 2

: : 129 2,. (gen. Hoplia) 3 2., rufipes,, obducta 3. 10 mm. 5 2,. djukini 8 mm. 5 4 5. aureola Genus Ectinohoplia Redtenbacher, 1868: 184. 10,,., 3. 2 1.. : Ectinohoplia sulphuriventris Redtenbacher, 1868: 184. : 32 (Loebl and Smetana, 2006) ( 2 ). :. : Kim (2001b). 57. Ectinohoplia rufipes (Motschulsky), 1860: 133 ( 6-50, 13-50, 19-50) Decamera rufipes Motschulsky, in Schrenk s Amur-Reise, II: 133, 1860. Hoplia rufipes: Heyden, 1887: 251; Bates, 1888: 372; Miwa and Chûjô, 1939: 73; Cho, 1957: 300; KZS, 1968: 137. Ectinohoplia rufipes: Niijima and Kinoshita, 1923: 80; Okamoto, 1924: 174; Mochizuki, 1935: 32; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 72; Cho, 1957: 300; 1963: 218; 1969: 683; Nomura, 1960: 57; Cho et al., 1968: 264; KZS, 1968: 137 ( ); Won and Woo,

130 1968: 379; KSPP, 1972: 206; Kim et al., 1975: 250; 1981b: 125; 1982a: 156; 1982b: 276; Kim, 1978: 358; Stebnicka, 1980: 261; Kim, 1981: 345; 1991: 66; 1995: 165; 2001b: 18; Nam and Kim, 1982: 129; Yoon et al., 1990b: 110; Park et al., 1993: 177; Park and Kim, 1993: 112; ESK and KSAE, 1994: 149; Kim et al., 1996b: 127; 1998: 170; 2002: 119. : (Cho, 1963 2), (KZS, 1968), (Cho, 1969 4), (KSPP, 1972), (Gu, 1973), (Kim et al., 1981). :.,., 1 2.,..,. 10 3.... Ad: ( ),.. BL: 7 10 mm. BW: 3.6 6 mm. : Murayama, 1931: 30; Medvedev, 1952: 155; Zhang, 1984: 92. :,,,,. :,,,. : NK-YG, HN, SK-GW, GG, SK- (6 ) 70 90 900. (774ex): [SWU] NK-1ex ( : 6.vii.1991). : 53ex (,,,,,,, : vi vii.1976 1998). : 457ex (,,,,,, : v viii.1962 1999). : 6ex (, : vi vii.1989 1991). : 14ex ( : v vi.1987 1996); 1ex ( : 19.vii.1999). : 1ex (, : 1.vii.1998); 57ex (,,,,,, : v viii.1976 1998). : 12ex (,,, : vi vii.1981 1997). : 6ex (vi viii.1959 1990). [NAAS] : 8ex ( : viii.1987). : 8ex (, : iv vii.1922 1991). : 2ex (, : vi.1975, 1998). : 23ex (vii x.1922 1985). [SNU] : 1ex ( : 27.vi.1997). : 75ex (,,,,,,, : iv x.1957 1998). : 7ex ( : vi xi.1984 1992). : 19ex (,, : v vii.1969 1997). [KU] NK-1ex ( : 10.v.1933). : 10ex (,,, : vi viii.1981 1994). : 171ex (,,,,,,, : iv viii.1932 1996). : 46ex (,,,,,, : vi viii.1960 1990). : 45ex (,,,, : v vii.1972 1993). : 65ex (vii viii.1955 1978). [EWU] : 2ex (, : v vi.1972, 1977); 3ex ( : viii.1979, 1980). : 54ex (,,

: : 131, : iv vi.1955 1991). : 2ex ( : v, vii.1988, 1993). : 1ex ( : 24.v.1981). : 1ex (11.vii.1979). [HSU] : 36ex ( : iv vi.1994 1997). : 676, 4 11. (iv- 6, v- 92, vi- 393, vii- 94, viii- 81, ix- 2, x- 6, xi- 2ex). : Saito, 1931: 473; 1941: 11; Nakayama and Okamoto, 1940: 198. 1. Ectinohoplia obducta (Motschulsky, 1857), Korean-Long-legged Scarab Beetle. Hoplia obducta Motschulsky, Etud. ent.: 33, 1857: Nomura, 1960: 57; KSPP, 1972: 206 ( ); Paik, 1984: 177 ( ); ESK and KSAE, 1994: 149. - Kim and Nam, 1982a: 156. :. : 6.5 11 mm. :,,,,. Genus Hoplia Illiger, 1803: 226. 1 2. 9 10,,. 6 7 1. 1 2. 2, 4. : Scarabaeus farinosus Linnaeus, 1761; 184 (Medvedev, 1952 ). : 250 ( 170 )( 2). :. : Kim (2001b).

132 58. Hoplia aureola (Pallas), 1871: 18 ( 6-51, 14-51, 19-51) Scarabaeus aureola Pallas, Icon. Ins. Ross., I: 18, pl. A, f. 10, 1871. Hoplia aureola: Niijima and Kinoshita, 1927: 29; Yoshino, 1935: 13; Mochizuki, 1935: 32; Mochizuki and Tsunekawa, 1937: 91; Haku, 1937: 120 (auleola); Miwa and Chûjô, 1939: 72; Cho, 1969: 684 ( ); KSPP, 1972: 207; Kim et al., 1975: 251; 1985: 105; Stebnicka, 1980: 260; Paik, 1984: 182 (accureolka); Kim et al., 1991e: 179; 1991e: 66; 1995: 170; ESK and KSAE, 1994: 149; Lie et al., 1997: 84. Hoplia (Euchromoplia) aureola: Kim, 2001b: 19. Hoplia aureula: Kim, 1978: 359; Kim and Nam, 1982a: 156; 1982b: 276; 1984: 328. Hoplia 12-punctata Motschulsky, in Schrenck s Amur-Reise, II: 133, 1860: Sawada, 1938: 12 (Syn. nov.); Heyden, 1887: 251. : (KSPP, 1972), (ESK and KSAE, 1994). :., 4, 3.... 5 4 5,. Ad: ( ),.. BL: 7 8 mm. BW: 3.7 4 mm. : Murayama, 1927: 12; 1931: 26; Medvedev, 1952: 158. : -,. - ( Nakayama and Okamoto, 1940). :,,,. :,,,. : NK-HB, HN, PB, PY,, PN,,, WB,, GG, SK-GW,,,,,,,, GS,,,,,,,,,,,,, (11 ),,,,,,,, CB,, GB,,, JB, JN, JJ. (244ex): [SWU] NK-18ex (,, : iv, v.1970 1985). : 1ex ( : 6.v.1989); 1ex ( : 8.vi.1985); 1ex ( : 28.v.2005). : 9ex (,,, : v, ix.1961 1993); 5ex ( : 13.vi.2009). : 1ex ( : 5.v.1998). : 1ex (1.viii.1972). [NAAS] : 1ex ( : 8.vi.1994). : 10ex ( : v.1922 1929). [SNU] : 1ex ( : 10.vi.1987); 1ex ( : 14.v.1979).

: : 133 : 5ex ( : 10.v.1987). [KU] : 1ex ( : 22.v.1977); 1ex ( : 27.vii.1958). : 122ex (,,,, : v vi.1934 1984). : 1ex ( : 8.vi.1974). : 9ex ( : v.1983). [EWU] : 1ex ( : 7.vi.1979); 3ex ( : 19.v.1985). : 29ex (,,, : v vi. 1959 1989). [HMNH-NK] : 6ex ( : 20 22.v.1985). : 13ex ( : 28.v.1970). : 2ex ( Sacheon-tong: 6 8.vi.1970). : 244, 4 11. (iv- 6, vi, v- 234, vii- 1, viii- 1, ix- 2ex). : Murayam, 1938: 259; Nakayama and Okamoto, 1940: 199. 59. Hoplia djukini Jacobson, 1914: 2 Hoplia djukini Jacobson, Russ. Emtom. Obozr. 14: 2, 1914: Stebnicka, 1980: 260; ESK and KSAE, 1994: 149 ( ). Hoplia (Decamera) djukini: Kim, 2001b: 20.,.,. 10 3...,.,. 5 2,. BL: 10 11 mm. : (Primorskij Kraj),. :. : NK-HN, 1, 11.vi.1965, Mroczkowski and Riedel (by Stebnicka, 1980). : Stebnicka(1980) Zoological Institute of the Polish Academy of Sciences (Warszawa) 1, Primorskij.. Hoplia 1..

134 Hoplia koreana Moser (Stett. Entom. Zeit., 81: 27, 1920) : Nigra, supra squamis viridi-mandallicis sat dense vestita. Capite granuloso-punctato, frontis punctis sandosis vel squamosis, clypeo antrorsum haud angustato, margine antico fortiter reflexo, truncato, angulis anticis rotundatis; antennis piceis, 9-articulatis; protho-race latiore quam longiore, lateribus ciliatis, curvatis, post medium levissime sinuatis, angulis anticis acutis, porrectis, angulis posticis fere rectangulis, prothoracis dorso elytrisque sat crebre squamosis and breviter erecte flavo-sandosis; pygidio corporeque infra dense viridi-vel coeruleo-argenteo-squamosis, pectore pallide sandoso-piloso, abdominis segmentis transversim sandosis; tibiis auticis tridentatis, unguibus anticis and mediis longitudine valde inaequalibus, unguibus posticis integris. -Long. 11 12 mm. Korea (Pu Ryong). C Rost leg. : Niijima and Kinoshita (1923: Seoul, 1927 ); Kim, 1978: 360 (Gnorimus subopacus: ); Won and Choi, 1968: 379; Yoon et al., 1979: 150; Kim et al., 1975: 251; 1982a: 156; 1982b: 276; Kim et al., 1991e: 179; 1995a: 165; 1998: 170; Miwa and Chûjô, 1939: 75; Cho, 1957: 300 ( ); 1969: 684; KZS, 1968: 137; ESK and KSAE, 1994: 149. Hoplia (= ) 1. Hoplia communis Waterhouse, 1875 Hoplia communis Waterhouse, Trans. Ent. Soc. Lond.: 100, 1875: Haku, 1937: 74 ( ); ESK and KSAE, 1994: 149 ( ) H. aureola. 2. Hoplia moerens Waterhouse, 1875 Hoplia moerens Waterhouse, Trans. Ent. Soc. Lond.: 100, 1875: Cho, 1969: 685 ( ); Kim and Kim, 1972b: 84; Kim, 1978: 360; Paik, 1984: 182 (H. morevs); ESK and KSAE, 1994: 149 H. aureola. 3. Hoplia squamacea White, 1844 Hoplia squamacea White, Ann. Mag. Nat. Hist., 14: 424, 1844: Miwa and Chûjô, 1939: 73; Cho, 1957:

: : 135 300; KZS, 1968: 138 (sauamacea, ); ESK and KSAE, 1994: 149,. Subfamily Melolonthinae Leach in Samouelle, 1819: 189. Melolonthidae Leach in Samouelle, 1819.,.. 8 10.,. 6,., (, spurs),.. 10,000, (Sabatinelli and Pontuale, 1998). 8 (Rhizotrogini) (Melolonthini) 2 11. 55. 20, 6 7. 30. : Melolontha Fabricius, 1775: 31. : 8 ( ) 28 ( )( 2/8 ). :? ( 12 ). : 10,000 ( )( 26 ). :. : Reitter (1902), Moser (1915a, 1919, 1920), Nijiima and Kinoshita (1923, 1927), Murayama (1954), Nomura (1973), Sabatinelli and Pontuale (1998), Kim (2001a, b). 1. 3. Rhizotrogini 2 3. Melolonthini 10 2. 10 3

136 9 8 3. 3 4 3 7 4... Apogonia. 5 5.. Brahmina 2 3 6 6., Sophrops, Holotrichia 7. 7 2. 5. Heptophylla 5. Hilyotrogus 8. Lasiopsis 9 9. 15 mm, Pseudosymmchia 18 mm. Miridiba 10. 7. 5 6. Polyphylla 3 2 1.2, 2. Melolontha Tribe Rhizotrogini Burmeister, 1855: 308. Rhizotrogidae Burmeister, 1855: 308.,. 9 10, 3. 3., 29. : Rhizotrogus Latreille, 1825: 371.

: : 137 :? ( 12 ). : 1,400 ( 29 ). :. : Reitter (1902), Moser (1915a, 1919, 1920), Nijiima and Kinoshita (1923, 1927), Murayama (1954), Kim (2001a, b). Genus Apogonia Kirby, 1818: 401. 5 15 mm,.,. 10 9.,. 2 3. 5 1 1 (Bezděk 2008), 2. : Apogonia gemellata Kirby, 1819: 401 (monotype). : 346 [ 35 (Bezdděkk, 2006)] ( 2). :. : Kim (2001b), Bezdděkk (2008), Sawada (1940). 1. 9 mm., cupreoviridis 8 mm. cribricollis 60. Apogonia cribricollis Burmeister, 1855: 256 ( 6-52, 14-52, 19-52) Apogonia cribricollis Burmeister, Handb. Ent., IV(2): 256, 1855: Murayama, 1954: 139; Cho, 1969: 667 ( ); Kim and Kim, 1974b: 108; Stebnicka, 1980: 262; Kim et al., 1991e: 67; 1997: 233; 1999: 129; 2004: 116; Kim, 2001b: 22; Park et Kim, 1993: 112; ESK and KSAE, 1994: 150. Apogonia amida Lewis, 1896 ( ); Morita, 1936: 851 ( ); ESK and KSAE, 1994: 150 (

138 ). Apogonia chinensis Moser, 1918 ( ): Niijima and Kinoshita, 1927: 16 ( ); Miwa and Chûjô, 1939: 62; Sawada, 1940: 277; Cho, 1957: 297; KZS, 1968: 137 ( ); ESK and KSAE, 1994: 150..,. 10, 3. 2,., 3 4.. 1. Ad:,..,. BL: 7 8.5 mm, BW: 4.3 5 mm. : Zhang, 1984: 88. :,,. :,,. : NK-HB, HN, PN, SK-GW,,,,,,,,,, CBN,,,, GB,,,,, JBN,,,. (64ex): [SWU] : 15ex (, : vi.1980, vii.1985); 1ex ( : 4.v.2003). : 1ex ( : 7.vii.1981). : 1ex ( : 26.vii.1979). : 1ex ( : 30.vi.1998); 4ex (,,, : vi viii.1986 1998). : 1ex (vii.1991). [NAAS] : 1ex ( : 16.x.1974). : 3ex ( : iv x.1923, 1977). : 1ex ( : 17.vi.2003); 7ex ( : v.1963 65). : 1ex ( : 20.viii.1996). : 1ex ( : 30.vii.1998); 1ex ( : 20.vii.1988). : 1ex ( : 25.vi.1994). [SNU] : 1ex ( : 12.ix.1993). : 3ex ( : v.1992); 4ex ( : v vi.1990 1992). : 2ex ( : viii.1979). [KU] : 1ex ( : 6.vi.1991); 1ex ( : 28.v.1982). : 1ex ( : 22.vi.1993). [EWU] : 1ex ( : 29.vii.1983). : 8ex (,,, : v vi.1959 1987). [HSU] : 2ex ( : v, vi.1994, 1995). 61. Apogonia cupreoviridis Kolbe, 1886: 193 ( 6-53, 14-53, 19-53) Apogonia cupreoviridis Kolbe, Arch. naturg. 52: 193, 1886: Bates, 1888: 373; Niijima and Kinoshita, 1923: 73; Maruta, 1929: 369; Miwa and Chûjô, 1939: 62; Mochizuki and Tsunekawa, 1937: 90;

: : 139 Murayama, 1954: 137; Cho, 1957: 298; 1963: 221; 1969: 666; Cho et al., 1968: 265 ( ); Stebnicka, 1980: 262; Yoon and Nam, 1979: 149; Kim et al., 1975: 240; 1981: 125; 1982a: 155; 1982b: 276; Kim, 1978; Kim et al., 1987b: 223; 1987c: 505; 1991b: 67; 1991e: 179; 1994b: 111; 1996b: 127; 1997: 233; 1999: 129; 2004: 116; Kim, 1993: 64; 1996c: 174; 2000: 132; 2001: 23; Park and Kim, 1993: 112; Park et al., 1993: 177; ESK and KSAE, 1994: 150; Lie et al., 1997: 60. Apogonia cupreoviridis fusania Kolbe, Arch. F. Naturf., 52: 193, 1886 (Fusan, Korea): Niijima and Kinoshita, 1923: 73; Haku, 1937: 121; Cho, 1957: 298. Apogonia nigroolivacea Heyden, 1887: Kalinina, 1989: 424; ESK and KSAE, 1994: 150; Bezděk, 2008: 72 (Syn. Nov.). : (Cho, 1963); nigroolivacea= (ESK and KSAE, 1994). : (, ).,,.,. 10, 3. 3... Ad:,..,.,.. BL: 8 11 mm, BW: 5 7 mm. : Murayama, 1931: 33; Medvedev, 1952: 108; Zhang, 1984: 87. :,, (Ryukyu). :,,,. : NK-HB Seisin (= ),,, HN,, PB, PY, PN,, WN, SK- 80 100 600. (544ex): [SWU] : 2ex (, : vi.1980, vii.1990); 1ex ( : 25.v.2001); 3ex (, : vi, vii.1993, 1999). : 62ex [,,,,,,,,, (6 ),,,, : iv xi.1971 1998]; 1ex ( : 4.vii.2007); 2ex ( : 29.v.2005, 17.vi.2008); 1ex ( : 28.v.2007). : 1ex ( : 17.vi.2003); 14ex (,,, : v vii.1986 1999). : 5ex (, : 29.vi 3.vii.1998); 13ex (,, : v vii.1983 1999). : 11ex (,,,,, : v viii.1982 1998). [NAAS] : 12ex ( : v, vi.1993 1996). : 28ex ( : iv xi.1923 1992). : 3ex ( : vii, viii.1996). : 2ex (, : vii.1988, 1993); : 14ex (,, : vi viii.1975 1994). [SNU] : 1ex ( : 20.vii.1997). : 135ex (,,,,,,,, : iv ix.1955 1997). : 3ex (,, : vii xi.1967, 1985, 1994). : 13ex (,, : v viii.1961

140 1998). : 5ex (iv vi.1967 1987). [KU] : 5ex (,, : v vii.1935, 1977, 1984). : 62ex (,,,,, : iv viii.1961 1996). : 5ex ( : vii.1988). : 2ex (, : vi.1989, 1990). : 8ex (,,, : v vii.1976 1993). : 5ex (v vii.1963 1978). [EWU] : 1ex ( : 5.vi.1977). : 20ex (,,, : v x.1959 1999). : 1ex ( : 20.v.1988). : 1ex ( : 15.viii.1971). : 1ex ( : 27.vi.1982). [HSU] : 1ex (9.vi.1997). [HMNH-NK] : 2ex ( : 22.vii.1975). : 1ex ( : 1.viii.1971); 2ex ( : 24.v.1970); 1ex ( : 8.vii.1977); 41ex ( : 13 15.vi.1985); 40ex (30.vi, 3.vii.1991). : 2ex ( : 3.vii.1988). : 1ex ( : 5.vii.1956). : 9ex ( : 4 7.viii.1975). : 1ex ( : 29.vii.1982). : 518, 4 11. (iv- 26, v- 160, vi- 206, vii- 90, viii- 26, ix- 4, xi- 4ex). : Murayama, 1954: 138 9; Kim and Kang, 1993: 415. Genus Brahmina Blanchardt, 1850: 140. 10.,.. Brenske (1892) B. heydeni Sophrops. Reitter (1902) 2, 20 Niijima Kinoshita (1923) 3. 6 5 1 8, 43. 3 5.,. 2.. : Melolontha cylindrica Gyllenhal, 1817. : 78 (Loebl and Smandana, 2006) ( 3 + 2 ). : -, -, -. : Murayama (1954), Kim (2001b). 1.. 2, excisiceps

: : 141 2... rubetra faldermanni 3 3.,.. intermedia. 4 4.,,.. 16 mm crenicollis,.. 16 mm darcis 62. Brahmina rubetra faldermanni (Murayama), 1954: 88 ( 6-54, 14-54, 20-54) Melolontha rubetra Faldermann, Mem. Acad. St. Petersb., 2: 376, 1935. Brahmina rubetra: Stebnicka, 1980: 264; ESK and KSAE, 1994: 150. Brahmina (Brahminella) rubetra: Loeble and Smetana, 2006: 212. Brahmina faldermanni Kraatz, Deut. Ent. Zeit., 1892: 309. Brahmina rubetra faldermanni (Murayama), Icons. Scarab. Manch. Kor., 1: 88, 1954: Cho, 1969: 670 ( ); Kim and Chang, 1987a: 104; Kim, 1993: 64; 2001b: 25. :, ( ).,,. 10, 3 6, 5...,...,.. Ad:. 2.5,.,. BL: 9 11 mm, BW: 5 6 mm.

142 : Zhang, 1984: 84 (B. faldermanni). :,. :,. : NK- : 1 ( : 22.vii.1922); 1ex ( : 9.viii.1959), SK-1ex ( : 24.vii.1986, Kim J.I.). (51ex): [SWU] : 1ex ( : 24.vii.1986, Kim J.I.). : 6ex ( : vi, vii.1990). [NAAS] : 1ex ( : 30.vii.1998). [HMNH-NK] : 2ex ( : 17.viii.1956); 1ex (13.ix.1980); 33ex ( : 16.vii.1975); 6ex (6 7.vii.1982); 1ex ( : 27.vii.1982). 63. Brahmina intermedia (Mannerheim), 1849: 238 Rhizotrogus intermedia Mannerheim, Bull. Soc. Nat. Hist. Moscou, 22: 238, 1849. Brahmina intermedia: Reitter, 1902: 180; Murayama, 1954: 90; Cho, 1957: 299; 1969: 670; KZS, 1968: 137 ( ); Stebnicka, 1980: 207; ESK and KSAE, 1994: 150; Lie et al., 1997: 63; Kim, 2001b: 27. Brahmina intermedia sibirica Brenske, Berl. Ent. Zeit., 37: 89,412, 1892 (E. Siberia): Niijima and Kinoshita, 1923: 55; Murayama, 1935: 20; 1937a: 35; Miwa and Chûjô, 1939: 66. :..,.,..,,..,,.. BL: 15 17 mm, BW: 7 9 mm. :,,. :. : NK-HB (2, viii.1922,, : Murayama, 1954). :.

: : 143 64. Brahmina darcis Reitter, 1902: 180 Brahmina darcis Reitter, Best.-Tab., 50: 180, 1902: Murayama, 1954: 95; Cho, 1969: 671 ( ); Stebnicka, 1980: 207; ESK and KSAE, 1994: 150; Kim, 2001b: 26. : (Kuku-noor). (Holotrichia).,.,.,,.,.,.,. BL: 16 17.5 mm, BW: 8 9 mm. :,,. :. : NK-PB (, 1, IX.1936, : Murayama, 1954). :. Brahmina 1. Brahmina crenicollis (Motschulsky), 1854 Ancylonycha crenicollis Motschulsky, Etud. Ent., III: 64, 1854: Reitter, 1902: 181; Niijima and Kinoshita, 1923: 55; Miwa and Chûjô, 1939: 66; Murayama, 1954: 97; Cho, 1957: 299; 1969: 671 ( ); KZS, 1968: 137; Stebnicka, 1980: 207; ESK and KSAE, 1994: 150; Kim, 2001b: 26; Loeble and Smetana, 2006: 212. : Korea (Reitter, 1902). :,.,,..,.,.,.,.

144. BL: 13 16mm, BW: 6.8 8mm. :. :. : Reitter,. 2. Brahmina excisiceps Moser, 1915 Brahmina excisiceps Moser, Deut. ent. Zeit.136, 1915 ( Ho-chan): Niijima and Kinoshita, 1923: 55 (, ); 1927: 22; Miwa and Chûjô, 1939: 66; Cho, 1957: 299; KZS, 1968: 137 ( ); Stebnicka, 1980: 208 (excissiceps Murayama); ESK and KSAE, 1994: 150; Kim, 2001b: 27. :. :. Genus Lasiopsis Erichson, 1848 9,. 2 1, 5 Stebnicka (1980) 1 1. Murayama (1941) (1954). 2. : Melolontha canica Zubokov, 1829. : 15 (Loebl and Smetana, 2006) [ 1 (?), 2 (?)]. :. : Murayama (1941, 1954), Stebnicka (1980), Kim (2001b). 1...

: : 145, 1, 2. 12 mm sahlbergi..,.. 14 mm manchuricus 65. Lasiopsis sahlbergi (Mannerheim), 1949: 239 ( 6-55, 14-55, 20-55) Rhizotrogus sahlbergi Mannerheim, Bull. Soc. Hist. Nat. Moscou, 22(1): 239, 1849. Lasiopsis sahlbergi: Murayama, 1954: 79; Cho, 1963: 221; 1969: 670; Cho et al., 1968: 265; Kim et al., 1974b: 108; 1975: 241; 1982a: 155; Yoon and Kim, 1981: 47; Kim, 1993: 64; 1995b: 139; 2001b: 28; 2002: 119; ESK and KSAE, 1994: 150; Loeble and Smetana, 2006: 211. Lasiopsis formosanus Niijima et Kinoshita, Res. Bull.Exp. Forest Hokkaido Univ., IV: 21,83, 1927; Murayama, 1937a: 34; 1954: 80 (Syn. nov.); Miwa and Chûjô, 1939: 66; Cho, 1957: 299; KZS, 1968: 137. : sahlbergi= (Cho, 1963), (Cho, 1969 4); formosanus= (KZS, 1968). : (Dauria)....,. 9 3.. 2, 3.. Ad:.,. LPMB= 2:1:3. BL: 11 12 mm, BW: 5.5 mm. :,,,. :,,,. : SK-GW, GS,,,, GN, (? ), JJ. (12ex): [SWU] : 1ex ( : 19.viii.2001). : 1ex ( : 31.viii. 1981).?: 1ex ( : 1.vii.1985). [EWU] : 1ex ( : 16.vi.1973). : 1ex (12.vi.1973). [HMNH- NK] : 3ex ( : 7.vii.1982); 1ex (3.vii.1991); 3ex ( : 3.vii.1988).

146 66. Lasiopsis manchuricus Murayama, 1941: 75 Lasiopsis manchuricus Murayama, Trans. Biol. Soc. Manch., 4(2): 75, 1941: Stebnicka, 1980: 264; ESK and KSAE, 1994: 150; Kim, 2001b: 28 ( ); Loeble and Smetana, 2006: 211 (manchurica). : (ESK and KSAE, 1994). : ( ).....,,. 9.,. 2.5., 4,. BL: 14 mm, BW: 6.5 mm. :,. :. : NK-PN (1, 13.ix.1970, Bielawaki and Mrocskowski: Stebnicka, 1980). :. Genus Pseudosymmchia Dalla Torre, 1913: 224. Metabolus Fairmaire, 1887. Ablotemus Paulsen and Smith, 2003. Paulsen and Smith (2003) Metabolus Fairmaire (1887, Coleoptera) ( ) Metabolus Bonaparte (1854, Aves: Monarchidae) ( ) Ablotemus Paulsen and Smith, M. tumidifrons Fairmaire. Smetana and Smith (2006) Pseudosymmchia. 1. : Symmachia chinensis Brenske, 1892: 780.

: : 147 : ( 1 ). : (?). : Paulsen and Smith (2003), Smetana and Smith (2006). 67. Pseudosymmchia impressifrons (Fairmaire), 1887: 316 ( 6-56, 14-56, 20-56) Metabolus impressifrons Fairmaire, Rev. d Ent., VI: 316, 1887: KSPP, 1972: 207; Stebnicka, 1980: 208; Kim, 2001b: 29. Pseudosymmchia impressifrons: Loeble and Smetana, 2006: 212. Metalobus impressifrons ( ): Kim et al., 1991e: 66; 1991e: 179; 1997: 233; 2004: 116; Kim, 1993: 65; ESK and KSAE, 1994: 150. Rhizotrogus niponicus Lewis, Ann. Mag. Nat. Hist., (6)14: 399, 1895 (Syn.): Arrow, 1913: 399; Okamoto, 1924: 174; Miwa and Chûjô, 1939: 67; Cho, 1957: 299; KZS, 1968: 137; Nomura, 1960 (Syn. nov.); KZS, 1968: 137. Miridiva koreana Niijima and Kinoshita, Bull. Exp. For., Hokkaido Univ., 2(2): 40, 1923 ( )(Syn.): Okamoto, 1924: 173; Maruta, 1929: 367; KZS, 1968: 137; Miwa and Chûjô, 1939: 67; Murayama, 1954: 83; Cho, 1957: 299; Nomura, 1960 (Syn. nov.); Cho et al., 1968: 264; Kim et al., 1972b: 197; 1976: 102; 1982b: 276; Nam and Kim, 1982: 129. Miridiba koreana ( ): Cho, 1969: 671 ( ); Kim et al., 1972b: 197; 1976: 102; 1982b: 276; Stebnicka, 1980: 265; Yoon and Kim, 1981: 47; Nam and Kim, 1982: 129. Miridiva coreana ( ): Eguchi, 1932: 51; ESK and KSAE, 1994: 150. : Rh. niponicus= (KZS, 1968); Miridiba koreana= (Cho, 1963 2); Metabolus impressifrons= (KSPP, 1972). : (Peiking). (Rhizotrogini) 9, 1, 1.,..,,,... Ad:,..,. BL: 10 15 mm, BW: 6 8 mm.

148 :,,. :,,,. : M. koreana: NK-HN, PB-, PN, WH, GW, SK-GW,,,, GS-,,,,,,, (17 ),,,, CB,,, GBN,, JN,,,,, JJ ; M. impressifrons: SK-GW,,,,, GS,,,,,,,,,,,,,,,,, CB,,,, GBN,,, JBN,,,,, JJ. (847ex): [SWU] : 1ex ( : 3.vii.1993). : 1ex ( : 10.viii.1980); 37ex [ (8 ), : vi x.1979 1998]; 14ex [ (4st.): vi viii.2003 2009]. : 1ex ( : 17.vi.2003); 1ex ( : 12.vii.1991). : 1ex ( : 10.vii.2000). : 4ex (,, : v x.1982 1998). : 1ex (27.vi.1994). [NAAS] : 180ex ( : iv x.1921 1996). : 2ex (vi, vii.1975, 1987). [SNU] : 2ex ( : ix.1993); 1ex ( : 24.vii.1974). : 359ex (,,,,,,,,, : iv x.1965 1997). : 7ex (,,, : vi ix.1989 1995). : 26ex (, : vi vii.1969 1997). : 2ex (13.vii.1993). [KU] : 4ex ( : vi vii.1979); 3ex ( : vi.1984 1994). : 23ex [, (3 ),,, : v vii.1934 1990]. 43ex (,, : vi.1961 1990). : 3ex ( : vi vii.1976). [EWU] : 1ex ( : 12.viii.1971). : 89ex (,,,,, : v vii.1955 1988). : 1ex ( : 6.vii. 1991). : 1ex (12.vii.1973). [HSU] : 1ex ( : 23.vi.1994). [HMNH-NK] : 1ex ( : 5.vi. 1991). : 1ex ( : 6 7.vii.1982); 1ex (26.v.1985); 2ex (6.vi, 3.vii.1991). : 25ex ( : 5.vii.1956). : 1ex ( : 17.vi.1985).?: 1ex (Mt. Tschonbonsan: 1.vii.1956); 2ex (Kuson and Tschondtsin: 1 29.vii.1956); 5ex (Chon-Ge-Charzon Wald: 12.viii.1956). : 725, 4 10. (iv- 7, v- 48, vi- 445, vii- 194, viii- 210, ix- 6, x- 4ex). : Murayama, 1935: 196; 1938a: 259 264; Saito, 1928: 12. Genus Miridiba Reitter 1902: 170. 1, Coca-Abia (2008) Holotrichia, Melolontha, Neodontocnema 3 8. Heyden (1887), Okamoto (1924) Morita (1936) 1 4,. (Hoikusha, 1956). : Rhizotrogus trichophorus Fairmaire, 1891. : 27 (Coca-Abia, 2008) ( 1 ).

: : 149 :. : Heyden (1887), Hoikusha (1956), Coca-Abia (2008). 68. Miridiba castanea (Waterhouse), 1875: 104 Holotrichia castanea Waterhouse, Trans. Ent. Soc., Lond.,: 104, 1875: Okamoto, 1924: 173 ( ); Morita, 1936: 860 ( - ); Miwa and Chûjô, 1939: 63; Cho, 1957: 298; Stebnicka, 1980: 208; KZS, 1968: 137. Lachnosterna (Holotrichia) castanea: Heyden, 1887: 251. Holotrichia (Pledina) castanea: Niijima and Kinoshita, 1923: 44; Lie et al., 1997: 66. Miridiba castanea: Cho, 1969: 672; Stebnicka, 1980: 208; Kobayashi, 1982a: 4 ( ); ESK and KSAE, 1994: 150; Kim, 2001b: 30. Miridiva castanea ( ): Murayama, 1954: 85; Cho, 1963: 218 ( ); Cho et al., 1968: 264. Neodontocnema castanea ( ): Kim et al., 1971: 160; KSPP, 1972: 208. : (Cho, 1969). : (Kawachi).,..,. 9, 3,.,.,., (, spurs), 5., 1. BL: 18 22 mm, BL: 10 11.5 mm. :,,,. :?,. : SK- ( ),, - (Kim and Kim: 1971). (3ex): [SWU] : 2 ( : 5.vi.2005). [EWU] : 1 ( : 8.vii.1965). Genus Sophrops Fairmaire, 1887: 106.,. 10 3.

150. 3. 2,. : Sophrops parviceps Fairmaire, Ann. Soc. ent. Belg., 1887: 106. : 38 (Loebl and Smetana, 2006) ( 2). : (? ). 1. 2.,. 13 mm striata.,. 12 mm heydeni 69. Sophrops heydeni (Brenske), 1982: 108 ( 6-57, 14-57, 20-57) Brahmina heydeni Brenske, Berl. Ent. Zeit., 37: 108, 1892. Sophrops heydeni: Reitter, 1902: 173; Niijima and Kinoshita, 1927: 25; Maruta, 1929: 368; Murayama, 1935: 4; 1937a: 34; 1954: 101; Mochizuki and Tsunekawa, 1937: 92; Miwa and Chûjô, 1939: 67; Cho, 1957: 299; 1963: 221; 1969: 672; Cho et al., 1968: 265; KZS, 1968: 137; Kim et al., 1972a: 84; 1974b: 108; 1982a: 155; 1982b: 276; 1985: 105; Kim 1978: 342; Yoon and Nam, 1979: 150; 1990b: 110; Stebnicka, 1980: 264; Kim et al., 1987b: 223; 1991: 66; 1996b: 127; 1999: 129; 2001: 132; 2004: 116; Kim, 1993: 64; 1995b: 139; 1996c: 174; 2001b: 31 ( ); Park et al., 1993: 178; Park and Kim, 1993: 113; ESK and KSAE, 1994: 150. : (Cho, 1963 10), (Kim, 1978 1). :.,,..,. 10,.. 2, 4.,. Ad:..., 1.2. BL: 10 11 mm,

: : 151 BW: 5 6 mm. : Zhang, 1984: 80. :,. :,,,. : NK-HB,, HN,, PB, PN,, WH, GW, SK- (, ) 60, 70 200. (153ex): [SWU] : 4ex (, : vii, viii.1995 2000). : 6ex (,,, : vii ix.1987 1996). : 10ex (,,,,,, : vii, viii.1976 2000). : 2ex ( : 29.vi; 2.vii.1998); 13ex (,,,,,, : vii, viii.1977 1998). : 6ex (,,, : vii, viii.1991 1998). [NAAS] NK-2ex ( : 10.ix.1921); 1ex ( : 25.vii.1924). : 17ex ( : v ix.1922 1996). : 1ex ( : 8.ix.1998). : 2ex ( : 2.vii.1998); 2ex ( : viii.1993). : 3ex ( : viii.1994 1998). [SNU] : 5ex (, : vii ix.1929, 1958 1996). : 11ex ( : vii.1969 1982). : 3ex (vii.1971). [KU] : 1 ( : vii.1928). : 32ex (,,,,, : vi viii.1974 1990). : 9ex (,, : vii, viii.1973 1990). : 3ex (, : vii.1977, 1984). : 5ex (, : vii viii.1973, 1976). : 1 ( : 3.viii.1955). [EWU] : 1ex ( : 29.vii.1983). : 11ex (,, : iv x.1954 1997). : 1ex ( : 1.viii.1966). [HMNH-NK] : 1ex ( : 10.vii.1977). : Nakayama and Okamoto, 1940: 200. 70. Sophrops striata (Brenske), 1892: 112 ( 6-58, 14-58, 20-58). Brahmina striata Brenske, Berl. Ent. Zeit., 37: 112, 1892. Sophrops striata: Niijima and Kinoshita, 1927: 24; Maruta, 1929: 368; Murayama, 1935: 4; 1937a: 34; 1954: 99; Miwa and Chûjô, 1939: 67; Cho, 1957: 299; 1969: 672 ( ); KZS, 1968: 137; Kim et al., 1972b: 197; 1974b: 108; 1975: 242; 1982a: 155; 1982b: 276; 1984a: 103; 1984b: 328; 1985: 105; Kim, 1978: 341; Shin, 1979: 144; Stebnicka, 1980: 265; Kim, 1981: 344; 1993: 64; 2000: 132; 2001b: 32; Kim et al., 1987a: 104; 1991e: 66; 1994b: 111; 1996b: 127; 1997: 233; 1998: 170; 2001: 132; 2002: 120; 2006: 36; Yoon et al., 1990b: 110; ESK and KSAE, 1994: 150. : (Kim and Nam, 1982b 4), (KZS, 1968). :.,,.,

152 2.. 10,,.,. 2, 4.,. Ad:..,.,. BL: 11.5 14 mm, BW: 5 6 mm. :,. :,,,. : NK-HN, PB ( ), PY, PN, WH,, GW, SK- (20 ) 70 90 1,100. (962ex): [SWU] : 10ex (,,,,, : iv vi.1984 2002); 2ex ( : 20.vi.2001). : 132ex [,,,,,, (7 ),,,,,, : iv viii.1971 2000). : 13ex (,,,, : iv vii.1994 1999). : 42ex (,,,,,,,,, : v vii.1986 1999); 1ex ( : 5.vi.2008). : 32ex (,,,, : iv vi.1987 1999). [NAAS] NK-2ex ( : vi.1923). : 1ex ( : viii.1987); 1ex ( : 31.v.1987). : 52ex (, : iv vi.1914 1996). : 7ex ( : v.1991); 1ex ( : 12.iv.1990). : 5ex ( : viii.1988). [SNU] : 1ex ( : 12.v.1992); 1ex ( : 11.vi.1988). : 265ex [,,,, (3 ),,,,, : iv ix.1957 1996). : 5ex (, : v vi.1990 1992); 2ex ( : 24.viii, 27.ix.2000). : 2ex ( : vi.1991). : 16ex (, : vi.1988 1997). : 3ex (vi.1988). [KU] NK-1ex ( : vii.1926). : 3ex (,, : vi.1979 1996). : 58ex (,,,,,,,,,, : iv vi.1961 1991). : 5ex (,,, : iv, v.1966 1986). : 9ex (,, : v, vi.1978 1990). : 50ex (,,,, : v, vi.1972 1993). [EWU] : 8ex (,,, : v viii.1965 1989). : 109ex (,,,,,,, : iv vi.1955 1999). : 2ex (, : vi, vii.1964, 1970). : 1ex ( : 23.vi.1964). : 2ex (, : v, viii. 1959, 1981). [HSU] : 118ex ( : iv vi.1994 1997). [HMNH-NK] : 1ex ( : 24.v.1970). : 1ex ( : 29.v.1970); 3ex (1.vi.1970); 2ex (26.v.1985); 1ex (13.vi.1991). : 716, 4 9. (iv- 43, v- 376, vi- 271, vii- 10, viii- 13, ix- 3ex). : Murayama, 1938: 259.

: : 153 Genus Holotrichia Hope, 1837: 100. Eretusa Reitter, 1902. Eotrichia Medvedev, 1951. 10,.,.. 3 15 8. 2, 2. 2 10. : Melolontha serrata Fabricius, 1972. : 125 (Loebl and Smetana, 2006) ( 10 : 2 ). :. : Arrow (1913), Heyden (1887), Bates (1888), Chang (1965), Kim (2000a, 2001b). 1.,.. 20 mm 2,... 1/2,., niponensis 2. 1 3 6 3....,. parallela, 4 4.,.,. koraiensis, 18 mm.. 5

154 5......,. picea., 1/2., 10,.., ernesti 6. 7 8 7.... diomphalia... oblita 8.....,.,, reticulata. 9 9..,.,,. kiotoensis...,. sichotana 71. Holotrichia niponensis (Lewis), 1985: 398 ( 6-59, 14-59, 20-59) Lachnosterna niponensis Lewis, Ann. Mag. Nat. Hist., (6)VI: 398, 1895. Lachnosterna nipponensis Murayama ( ): Stebnicka, 1980: 207. Holotrichia niponensis: Niijima and Kinoshita, 1923: 54; Miwa and Chûjô, 1939: 64; Cho, 1957: 298; KZS, 1968: 137; Kim et al., 1991e: 66; 2001: 132; 2005: 79; ESK and KSAE, 1994: 150 ( );

: : 155 Kim, 2000a: 15; 2001b: 36. Holotrichia titanis Reitter, Verh. Nat. Ver. Brünn, XL, Best.-Tab., 50: 178, 1901 (Syn.: ): Niijima and Kinoshita, 1927: 18; Maruta, 1929: 368; Kamijo, 1932: 21; Miwa and Chûjô, 1939: 65; Murayama, 1954: 111; Cho, 1957: 299; 1963: 218; 1969: 668; Cho et al., 1968: 264; KZS, 1968: 137; Kim and Kim, 1972b: 84; 1977: 130; 1982b: 276; 1984a: 103; 1984b: 328; Nam and Kim, 1982: 129; Yoon and Nam, 1990b: 110; Park et al., 1993: 178; Lie et al., 1997: 66. Lachnosterna titanis: Nomura, 1960: 65 (Syn. nov.); KSPP, 1972: 207; Kim, 1978: 337; Yoon and Nam, 1979: 149; Kim and Nam, 1982b: 276. Eotrichia titanis: Yoon and Nam, 1979: 149; Stebnicka, 1980: 270; ESK and KSAE, 1994: 150; Kim, 1995b: 139; Kim and Kim, 1996b: 127. : niponensis= (KZS, 1968); titanis= (Cho, 1963), (KZS, 1968 12), (Cho, 1969 1), (ESK and KSAE, 1994). : Fusan (= ). (Nagasaki, Goto Islands)...,.,. 10, 3...,,.. Ad:.., ( ).,. BL: 17.5 22.5 mm, BW: 10 12 mm. : Murayama, 1931: 45 (H. titanis); Medvedev, 1952: 111 (H. titanis). :,,, (Kyushu). :,,,. : H. titanis=nk-hb, HN,, PB, PN ( ), ( ), PY,, WH,, SK- (18 ) 35 ; H. niponensis=nk-pn, SK-GW,,,,,,,,,,,,,,,,,,, JB,,, JJ. (216ex): [SWU] : 1 ( : 9.viii.1935). : 1ex ( : 12.vi.1997); 1ex ( : 28.v. 1987). : 32ex (,,,,,,, : iv viii.1961 2000); 1ex ( : 28.iv. 2008). : 3ex (,, : v vii.1991 1999). : 2ex ( : v.1993). : 2ex (v, vii.1994, 1997). [NAAS] : 5ex (, : iii v.1927 1993). [SNU] : 40ex (,,, : iii vi.1967 1991). : 1ex ( : 19.vi.1992). : 1ex ( : 18.v.1991). : 5ex (iv, v.1985 1995). [KU] : 4ex (,, : vi, v.1977 1995). : 30ex (,, : iv vii.1931, 1957 1995). : 2ex ( : v.1986). : 3ex (, : v, vii.1972, 1985). : 6ex (v viii.1955 1989).

156 [EWU] : 1ex ( : 31.v.1983). : 74ex (,,, : iv ix.1956 1987). : 1ex (8.v.1965). : 199, 3 9. (iii- 4, iv- 58, v- 96, vi- 25, vii- 7, viii- 6, ix- 3ex). : Murayama, 1935: 196; 1938a: 259 264; Kim and Kang, 1993: 415 (H. titanis). 72. Holotrichia parallela (Motschulsky), 1854: 64 ( 6-60, 7-60, 14-60, 20-60) Ancylonycha parallela Motschulsky, Etud. Ent. III: 64, 1854. Lachnosterna parallela: Kolbe, 1886: 193; Heyden, 1887: 251. Holotrichia parallela: Miwa and Chûjô, 1939: 64; Cho, 1957: 298; KZS, 1968: 137; Stebnicka, 1980: 297; Kim et al., 1991e: 67; 2002: 120; 2004: 116; ESK and KSAE, 1994: 150; Lie et al., 1997: 67; Kim, 2001b: 37. Lachnosterna morosa Waterhouse, Trans. Ent. Soc. Lond.: 104, 1875 (Japan, China): Heyden, 1887: 251; Bates, 1888: 373; Nomura, 1960: 65 (Syn. nov.); Kim, 1978: 336; Yoon and Nam, 1979: 149; Park and Cho, 1986: 127. Holotrichia morosa: Niijima and Kinoshita, 1923: 49; Okamoto, 1924: 17; Maruta, 1929: 368; Doi, 1935: 3; Kamijo, 1936: 86; Morita, 1936: 861; Miwa and Chûjô, 1939: 64; Murayama, 1954: 104; Cho, 1957: 298; 1969: 667; Cho et al., 1968: 264; Won and Choi, 1968: 379; KZS, 1968: 137 ( ); Hyun and Woo, 1969: 193; KSPP, 1972: 207; Kim and Kim, 1974b: 108; 1975: 240; 1982a: 155; 1982b: 276; Nam and Kim, 1982: 129; Yoon et al., 1986: 160; 1990a: 140; Park and Kim, 1986: 127; Kim et al., 1987b: 223; 1987c: 505; 1989: 176; 1991e: 67; 1991b: 179; 1997: 233; 1998: 170; 1999: 129; 2005: 79; Kim, 1993: 65; Park et al., 1993: 178; Park and Kim, 1993: 112; ESK and KSAE, 1994: 150; Kim, 2000a: 15. Lachnosterna inelegans Lewis, Ann. Mag. Nat. Hist. (6)VI: 396, 1895 (Japan, Fusan= )(Syn. nov.: Arrow, 1913). Holotrichia inelegans: Stebnicka, 1980: 269. : parallela= (KZS, 1968 1); morosa= (Cho, 1963; Cho et al., 1968), (KSPP, 1972). :...,. 10, 3. 1,

: : 157.,,. Ad:,.,.. 2. BL: 17 22 mm, BW: 9 11 mm. : Murayama, 1931: 41; Medvedev, 1952: 109; Zhang, 1984: 76. :,,,,,,,. :,,,,,. :,,,. : morosa=nk-yg, PB,, HN, GW,, SK- (8 ) 70 100 ; parallela=nk-pn, GW, SK- 90 130 1,500. (1,228ex): [SWU] : 18ex (,, : vi viii.1983 1999); 2ex ( : 19.viii.2001); 1ex ( : 6.vii.2006). : 115ex (,,,,,,,,,,,,,,,, : vi x.1977 2003). : 1ex (, : 17.vi.2003); 38ex (,,,,,,,, : vii viii.1971 1999); 3ex ( : 23.vi.2005). : 7ex ( : 29.vi 3.vii.1998); 38ex (,,,,,,,,,,,,,,,, : vi viii.1982 2000). : 84ex (,,,,,,,,,,,,,, : vi viii.1981 1998); 1ex ( : 27.vii.2002). [NAAS] : 3ex ( : vii.1993); 1ex ( : 21.ix.1992). : 173ex (,, : v ix.1921 1997). : 30ex (,,,, : vi viii.1987 1996). : 31ex (,,, : vi ix.1991 1993). : 52ex (,,,,,,,, : vi xi.1922, 1975 1991). : 220ex (vi ix.1973 1998). [SNU] : 6ex (, : vi vii.1988 1997). : 85ex (,,,,,, : vi ix.1967 1998). : 3ex (,, : vii viii.1970 1983). : 19ex (,, : vii, viii.1990, 1996). : 89ex (,,, : v ix.1968 1996). : 25ex (vii.1965 1997). [KU] : 5ex (, : vi.1925, 1994). : 34ex (,,, : iv viii.1955 1990). : 34ex (,, : vii viii.1960, 1973 1979). : 13ex (,, : vii ix.1964 1979)., : 20ex (,,,,, : vi viii.1933 1990). [EWU] : 1ex ( : 5.vi.1977). : 29ex (,,, : v xi.1960 1993). : 4ex (, : vii viii.1969, 1971). : 7ex (,,,, : vi vii.1964 1972). : 12ex (,,, : vii.1969 1991). : 1ex (13.vii.1979). [HSU] : 23ex ( : v vii.1994 1997). [HMNH-NK] : 9ex ( : 5 25.vii.1956). : 2ex ( : 4 6.viii.1975); 1ex (17.vi.1985); 1ex (13.vi.1991).?: 1ex (Tschon-bon-san: 3.ix.1956); 1ex (Tschondtsin: 29.vii.1956); 1ex (Vekki: 16.ix.1956). : 825, 4 10. (iv- 2, v- 18, vi- 130, vii- 417, viii- 214, ix- 36, x- 8ex). : Eguchi, 1932: 53; Murayama 1938: 259; Nakayama and Okamoto, 1940: 199; Saito 1941:

158 11; Kim and Hyun, 1988: 21 27 (morosa and diomphalia); Kim, 1990: 222 229 (diomphalia); Kim and Kang, 1993: 415. 73. Holotrichia koraiensis Murayama, 1937a: 37 ( 7-61, 14-61, 20-61) Holotrichia koraiensis Murayama, J. Chosen Nat. Hist. Soc. 22: 37, 1937a: Miwa and Chûjô, 1939: 63; Murayama, 1954: 116; Cho, 1957: 298; 1969: 669; KZS, 1968: 137 ( ); Stebnicka, 1980: 208; Kim et al., 1987c: 505; 1998: 170; Kim, 1993: 65; 1995b: 164; 2000a: 16; 2001b: 38; ESK and KSAE, 1994: 150. : (,, )..,...,.. 1,.,. Ad:,. ( )., (, lobes). BL: 17 20 mm, BW: 8.8 10.5 mm. :,, (Is. Tsushima). :,,,. : NK-HB, HN ( ), SK-,,,,,,,, GG,,,,,,,, GB,, JB, JN, JJ. (57ex): [SWU] : 4ex ( : vi vii.1970 1979); 3ex ( : 23.vi.1996); 1ex (Mt. : 24.vii.1991); 1ex ( : 16.vi.1993); 1 ( : 30.v.1999); 1ex (, : 21.vii.1998). : 4ex ( : vi.1996). : 1ex ( : 12.v.1986); 1ex ( : 10.v.1997). : 1ex ( : 25.v. 1993). : 1ex (7.vi.1972). [SNU] : 2ex ( : ix.1988). [KU] : 1ex ( : 13.vi.1988); 3ex ( : vi vii.1979 1991). : 2ex ( : vi.1934); 1ex ( : 24.v.198:8). : 6ex ( : 6.vi.1981); 2ex ( : 14.vi.1996).?: 1ex (Heihoku: 15.viii.1936). [EWU] : 1ex ( : 24. vi.1991). : 1ex ( : 2.v.1978). [HMNH-NK] : 11ex ( : 2 5.vi.1991). : 3ex ( : 23, 28.v.1991). : 4ex ( : 12.vi.1991).

: : 159 74. Holotrichia picea (Waterhouse), 1875: 102 ( 7-62, 14-62, 20-62) Lachnosterna picea Waterhouse, Trans. Ent. Soc., Lond.: 102, 1875; Heyden, 1887: 252. Holotrichia picea: Heyden, 1889: 660; Haku, 1935: 57; Miwa and Chûjô, 1939: 65; Murayama, 1954: 114; Cho, 1957: 298; 1963: 218; 1969: 668; KZS, 1968: 137 ( ); Shin and Choo, 1977: 90; Stebnicka, 1980: 268; Kim et al., 1987c: 505; 1991e: 66; 2005: 79; 2006: 36; ESK and KSAE, 1994: 150; Lie et al., 1997: 68; Kim, 2000: 132; 2000a: 16; 2001b: 39. : (, Cho, 1963). : (Ngasaki, Hakodadi).,..,..,..,. Ad:.,.,,.,. BL: 15 18 mm, BW: 8.5 9.5 mm. :,,,. :,,,. : NK-HN ( ), PY,, PN, SK-GW,, GS, (, ),, (5 ), CB, CN, GB,,, GN, JN,, JJ,,. (23ex): [SWU] : 1ex ( : 9.viii.1976). : 1ex ( : 26.vi.1987); 2ex ( : v.1967, 1997). : 1ex ( : 4.viii.1973). : 2ex ( : 29.v.1999); 1ex ( : 21.vi.1997). : 1ex ( : 9.vii.1978). [SNU] : 1ex ( : 17.vi.1969). : 1ex ( : 23.xi.1985). [KU] : 1ex ( : 18.v.1991). : 1ex ( : 3.v.1978). [EWU] : 4ex ( : vi, vii.1979, 1983). : 5ex (,,,, : v vii.1956 1971). : 1ex ( : 7.vi.1969). 75. Holotrichia ernesti Reitter, 1902: 177 ( 7-63, 14-63, 20-63) Holotrichia ernesti Reitter, Verh. Nat. Ver. Brünn. (Best-Tab.), 50: 177, 1902: Murayama, 1954: 120; Cho, 1969: 669; Stebnicka, 1980: 266; Kim et al., 1991e: 67; Kim, 1993: 65; 2000: 132; 2000a: 16; 2001b: 40;

160 Park and Kim, 1993: 112; ESK and KSAE, 1994: 150 ( ); Lie et al., 1997: 70. : (Cho, 1969). :., 10., 1/2.,..,. Ad:,..,,,.,.. BL: 16 19.5 mm, BW: 9 10 mm. :,,,. :,,,. : NK-HB,,, HN,, PN, WH, SK-GW,, GB,, GN, JB, JJ. (24ex): [SWU] : 2ex ( : 6.v.1973); 1 ( : 5.viii.1961). : 11 4 ( : 29.v.1999); 2 ( : 20.vi.1997); 2ex ( : 29.v.1999; 26.vii.2001). [EWU] : 1ex (8.vii. 1965). [HMNH-NK] 1ex (Tchon-bon-san?: 3.ix.1956). : Murayama (1954) 1 1. 76. Holotrichia diomphalia (Bates), 1888: 373 ( 7-64, 14-64, 20-64) Lachnosterna diomphalia Bates, Proc. Zool. Soc. Lond.: 373, 1888. Holotrichia diomphalia: Niijima and Kinoshita, 1923: 50; Okamoto, 1924: 174; Maruta, 1929: 368; Kamijo, 1932: 21; Eguchi, 1932: 52; Cho, 1934: 75; Haku, 1935: 57; Masaki, 1936: 260; Mochizuki, 1935: 32; 1936: 211; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 63; Mori and Cho, 1940: 13; Cho 1947: 65; 1963: 218; 1969: 667 (dimorphalia); Murayama, 1954: 106; Kim, 1956: 9; 1958: 98 ( ); Cho et al., 1967: 197; 1968: 264; Won and Choi, 1968: 379; KZS, 1968: 137; Kim et al., 1974b: 108; 1982a: 155; 1984b: 328; Stebnicka, 1980: 265; Kim et al., 1987c 505; 1991e: 66; 1994b: 111; 1996b: 127; 1997: 233; 2004: 116; Kim, 1993: 65; 1994a: 214; 1995b: 139; 1999: 129; 2000: 132; 2000a:

: : 161 16; 2001b: 40; Park et al., 1993: 178 (diompharia); Park and Kim, 1993: 112; ESK and KSAE, 1994: 150; Lie et al., 1997: 72. Lachnosterna niponensis Lewis, 1895 ( ); Kim and Yoo, 1987b: 505. : (Cho, 1947), ( ) (Cho, 1963 2). : ( )....,..,.. Ad:.,... ( ),,,..,. BL: 16 21 mm, BW: 8 10 mm. : Murayama, 1931: 38; Medvedev, 1952: 110; Zhang, 1984: 78. :,,,,,,. :,,,,. :,,,. : NK-HB, HN, WB, GW,, SK- 100 130 3,000. (2,456ex): [SWU] 115ex (,,,,,,,,,,,,, : v viii.1983 2002); 5ex ( : 25.vii.2000); 10ex ( : 18 22.vi.2001); 2ex ( / : 10/20.vii.2009); 1ex ( : 22.vi.2005); 1ex ( : 13.v.2006); 1ex ( : 17.vii.2002). : 224ex [,,,,,,,,, (7 ),,,,,, : iv ix.1977 2001]; 5ex ( : 25.vi.2005); 4ex [ (,, ): v.2009]; 1ex ( : 17.v.2006). : 1ex ( : 17.vi.2003); 47ex (,,,,,, : iv vii.1981 1999); 1ex ( : 6.vi.2006). : 9ex ( : 30.vi.1998); 3ex ( : 29.vi, 2.vii.1998); 217ex (,,,,,,,,, : v ix.1977 1998). : 27ex (,,,,,,,, : v viii. 1973 1997). : 26ex (v vii.1972 1997); 2ex ( : 16.viii.2009). : 3ex (10.vii.2002, 28.v. 2005, 4.vi.2006). [NAAS] NK-1ex ( : 5.vi.1923). : 1ex ( : 13.vi.1996); 3ex ( : viii. 1987). : 74ex (,,, : iii viii.1921 1996). : 11ex (,, : iv ix.1990 1997). : 5ex (,, : vii viii.1983 1993). : 33ex (,,,,, : vi viii.1922 1995). : 8ex (iv vii.1919 1998). [SNU] : 23ex (,,

162,, : v vii.1974 1998). : 378ex (,,,,,,,,,,,,,, : iv x.1957 1997). : 6ex (,,, : vi vii.1983 1995). : 4ex ( : vi.1987). : 86ex (,,,, : v viii.1969 1997). : 31ex (iv viii.1985 1997). [KU] NK-1ex ( : 16.vii.1930); 1ex ( : 20.vii.1932). : 45ex (,,,,, : vi vii.1931 1995). : 218ex (,,,,,,,,,,, : iv ix.1913 1993). : 21ex (,,, : v vii.1957 1985). : 19ex (,,,,,, : iv vii.1971 1996). : 66ex (,,,,,, : v viii.1972 1993). : 11ex (v viii.1955 1990). [EWU] NK-3ex ( : 28.v.1965). : 16ex (,,,,,, : v viii.1971 1991). : 286ex (,,,,,,, : iv x. 1957 1993). : 13ex (,,,, : v vii.1956 1971). : 13ex (,,,,,,, : v vii.1966 1999). : 26ex (,,,,, : iv ix.1960 1992). : 7ex (v vii.1965 1983). [HSU] : 246ex ( : iv vi.1994 97). [HMNH-NK] : 1ex ( : 22.vii.1975). : 1ex ( : 12.ix.1980); 6ex (20 22.v.1985); 4ex (23.v.1991); 30ex (27, 28.v.1991); 1ex (6.vii.1991). : 2ex (, : 30.vi.1991); 11ex ( : 17.v.1985); 17ex ( : 24, 30.v.1985). : 1ex ( : 12.vii.1977); 3ex (26.v.); 1ex (17.vi.1985); 6ex (10 13.vi.1991).?: 2ex (Tschon-bon-san: 1.vii.); 9ex (3.ix.1956); (Tschondtsin?: 29.vii.1956). : 1,806, 3 10. (iii- 11, iv- 81, v- 754, vi- 629, vii- 234, viii- 47, ix- 21, x- 1ex). : Okamoto, 1924: 174; 1940: 198; Saito, 1928: 12; Eguchi, 1932: 52; Nakayama, 1934: 80; Murayama, 1934a: 42 46; 1935a: 196; 1935b: 329; 1936: 1 6; 1938: 259 264; Kim and Hyun, 1988: 21 27; Kim, 1990: 222 229; Kim and Kang, 1993: 415. : 1. 77. Holotrichia oblita (Faldermann), 1835: 459 ( 7-65, 14-65) Ancylonycha oblita Faldermann, Mem. Acad. Sci. St. Petersb., 2: 459, 1835. Holotrichia oblita: Stebnicka, 1980: 268; ESK and KSAE, 1994: 150 (H. oblita Stebnicka- : ); Kim, 2000a: 17; 2001b: 42...... Ad: Chang (1965).. BL: 16 22 mm.

: : 163 : Zhang, 1984: 77 ( ). :,,,. :,. : NK-HN, PB PY,,, WH,, GW SK-GG, GB. (2 ): [SWU] : 1 ( : 28.iv.1959). : 1 ( : 5.v.1985). : Zhang (1984), 1 2 1. 3 7 4,., ( ). 78. Holotrichia reticulata Murayama, 1941: 39 ( 7-66, 14-66, 20-66) Holotrichia reticulata Murayama, Annot. Zool., Japan, 20(1): 39, 1941: Murayama, 1954: 115; Cho, 1963: 218; 1969: 669; Cho et al., 1968: 264; KZS, 1968: 137; Stebnicka, 1980: 208; ESK and KSAE, 1994: 150 ( ); Kim, 2000a: 17; 2001: 43. : (Cho, 1963). : ( )....,.,,. Ad:.,.,. 2, 2.,. BL: 17 21 mm, BW: 8.5 11 mm. :. :,. : SK-1 ( : 29.vi.1923, coll. Okamoto and Kurisué). (20ex): [SWU] : 1ex ( : 6.vi.1985). : 10ex ( : iv, v.1984 1997); 3ex (27 29.iv.2003). [SNU] : 4ex ( : iv, v.1985 1995). [KU] : 1ex ( : 3.v.1978). [EWU] : 1ex (27.v.1983).

164 : Holotrichia Microtrichia. Nomura (1970) Pollaplonyx Bunbunius H. reticulata.. Holotrichia 1. Holotrichia kiotoensis Brenske, 1894 Holotrichia kiotoensis Brenske, Mem. Soc. Ent. Belg. II: 19, 68, 1894 (Japan): Yoshino, 1935: 14; Murayama, 1937a: 35; 1954: 110; Miwa and Chûjô, 1939: 64; Cho, 1957: 298 (kiotonsis); 1969: 668; Won and Choi, 1968: 379; KZS, 1968: 137 ( ); Stebnicka, 1980: 208; Lee et al., 1994: 147; ESK and KSAE, 1994: 150; Kim, 2001: 43; Lachnosterna kiotoensis: KSPP, 1972: 207. Lachnosterna inelegans Lewis, Ann. Mag. Nat. Hist. (6)VI: 396, 1895 (Pusan)(Syn.: Miwa and Chûjô, 1939)./Holotrichia inelegans: Stebnicka, 1980: 269. Holotrichia diomphalia ( ): Kim and Kim, 1972a: 83 (kiotensis); 1977: 130; Park and Park, 1987: 673; Kim, 1987b: 224; 1987c: 505; 1991e: 67; 1997: 233; Park and Kim, 1993: 112; Kim et al., 2005: 79. : GN- (Lewis, 1895), JN- (Yoshino, 1935), (Murayama, 1937), (Won and Choi, 1968), GG-,,,, GN- 6.,..,,. Murayama (1954) (H. diomphalia). Ad:.. :,,?,. :,. : Lewis (1894). Yoshino (1935)..

: : 165 2. Holotrichia sichotana Brenske, 1896 Holotrichia sichotana Brenske, Berl. Ent. Zeit.: 343, 1896 (Wladiwostock): Niijima and Kinoshita, 1927: 17 ( ); Maruta, 1929: 368 ( ); Murayama, 1935: 4; 1937a: 35; 1954: 118; Miwa and Chûjô, 1939: 65; Cho, 1957: 298; 1969: 669; KZS, 1968 ( ): 137; Stebnicka, 1980: 208; Kim and Kim, 1982a: 155; ESK and KSAE, 1994: 150; Kim et al., 1991e: 66; 1998: 170. : ( ),,,,,,,,, ( Murayama, 1935). ( ),.,. Ad: Chang (1965) (H. koraiensis). :,,,?. :,,,. : Murayama (1935) (1954).. Holotrichia (= ) 1. Holotrichia convexopyga Moser, 1912 Holotrichia convexopyga Moser, Ann. Soc. ent. Belg., 435, 1912: Chang, 1965: 37; ESK and KSAE, 1994: 150 ( ); Lie et al., 1997: 70 ( ); Kim et al., 1999: 129 (H. diomphalia ). 2. H. novila O. Park and Han, 1992: 138 ( ).

166 Genus Heptophylla Motschulsky, 1858: 32. 7, 5,,.,. ( ).. : Heptophylla picea Motschulsky, 1858. : 30 [ 10 (Loebl and Smetana, 2006)] ( 1 ). :,. : Kim (2001b). 79. Heptophylla picea Motschulsky, 1857: 33 ( 7-67, 14-67, 20-67) Heptophylla picea Motschulsky, Etud. Ent., VI: 33, 1857: Miwa and Chûjô, 1939: 69; Kim, 1958: 98; Cho, 1957: 300; Cho et al., 1967: 197; KZS: 137 ( ); Hyun and Woo, 1969: 193; Won and Choi, 1968: 379; Kim and Kim, 1972a: 160; 1972b: 84; Shin 1979: 144; Stebnicka, 1980: 207; Nam and Kim, 1982: 129; Park et al., 1986: 128; 1993: 178; Kim et al., 1987a: 104; 1998: 170; ESK and KSAE, 1994: 150; Kim, 2001b: 45. : ( ) (Cho et al., 1967 2). :.,.,.,. 10 6.5 2, 5. 4,. Ad:,.,., 2 ( ).. BL: 11.5 15 mm, BW: 5 8 mm. :,,.

: : 167 :,. : SK-GW,,, GS,,, CN, GB, GN, JB, JN. (12ex): [SWU] : 3ex (, : v vii.1999); 1ex ( : 8.viii.1993); 1ex ( : 5.viii. 2005); 2ex ( : 20, 26.vii.2005); 1ex ( : 30.vii.2009). : 1ex ( : 19.vii.1999). : 1ex ( : 30.vi.2003); 2ex (, : 26.v.2003). : Leal, W.S. et al., 1996; Kakizaki et al., 1998: 5 10; 2000: 44 46. Genus Hilyotrogus Fairmaire 1886: 325. Melichrus Brenske, 1892. 10 5 2, 4 5... 1. : Hilyotrogus unguicularis Fairmaire, 1886. : 27 (Loebl and Smetana, 2006) ( 1 ). :?. : Fairmaire (1886), Murayama (1954). 80. Hilyotrogus bicoloreus (Heyden), 1887: 265 ( 7-68, 14-68, 20-68) Lachnosterna (Holotrichia) bicoloreus Heyden, Horae Soc. Ent. Ross., 21: 265, 1887. Hilyotrogus bicoloreus: Reitter, 1902: 258; Niijima and Kinoshita, 1923: 59; Murayama, 1937a: 35; 1938: 14; 1954: 127; Nagaoka, 1938: 25; Mori and Cho, 1940: 13; Miwa and Chûjô, 1939: 68; Cho, 1947: 65; 1957: 300; 1963: 221 ( ); 1969: 673; Cho et al., 1968: 265; KZS, 1968: 137; KSPP, 1972: 207 (bicolorens); Kim et al., 1974b: 108; 1976: 102; Kim, 1978: 344; Yoon et al., 1979: 150; 1981: 47; 1990b: 110; Stebnicka, 1980: 261; Kim, 1981: 344; 1993: 64; 1995b: 164; 1996c: 174; 2001b: 45; Nam and Kim, 1983: 129; Park and Park, 1987: 673; Park et al., 1993: 178; Park and Kim, 1993: 112; ESK and KSAE, 1994: 150; Kim et al., 1994b: 111; 1996b: 127; 1997: 233; 1998: 170; 1999: 129; 2000: 132; 2001: 132; 2002: 120; 2004: 116; Lie et al., 1997: 60.

168 : (Cho, 1947); (KSPP, 1972). :.,.,,.,. 2.,.,. Ad:. 2,.. 1.5.,. BL: 15 18 mm, BW: 8 8.5 mm. :,,. :,,,. : NK-HB, HN,,,, PN,, GW, SK- 70 600. (507ex): [SWU] : 99ex (,,,,,,, : vi viii. 1975 2000); 5ex ( / : 10/20.vii.2009). : 21ex (,,,,,,,, : vi x.1980 1998). : 22ex (,,,,,,, : vii.1987 2000). : 22ex ( : 29.vi 3.vii.1998); 33ex (,,, : vi ix. 1985 1998). : 49ex (,,,,, : vi viii.1981 1998). [NAAS] : 2ex (, : viii, ix.1923, 1987). : 4ex (,, : vii viii.1991 1998). : 7ex (,, Mt., : vii ix.1922, 1988 1998). : 11ex (,, : vi viii.1922 1995). [SNU] : 1ex ( : 17.vii.1962); 1ex ( : 12.ix.1997). : 7ex (,, : vii ix.1982 1998). : 4ex (,, : vi viii.1983 1997). : 57ex (,, : vi viii.1968 1998). [KU] NK-1ex ( : 16.xi.1930). : 33ex (,,, : vii viii.1958 1992). : 14ex (,,, : vi ix.1931 1985). : 11ex ( : vii ix. 1973 1979). : 32ex (,,,,, : vi vii.1967 1990). : 6ex (,, : vii viii.1974 1985). : 4ex (vii viii.1931 1974). [EWU] : 7ex (,,, : viii.1955 1983). : 2ex (, : vii.1956, 1965). : 4ex (, : vii, viii.1969 1987). : 4ex ( : 29.vii.1983). [HSU] : 8ex ( : v vi.1996, 1997). [HMNH- NK] : 1ex ( : 17.viii.1956); 4ex (12 14.ix.1980); 3ex (13 15.vii.1982); 1ex (5.vii.1991). : 1ex ( : 29.ix.1978). : 5ex ( : 9 12.vii.1977). : 424, 5 10.(v- 2, vi- 83, vii- 222, viii- 98, ix- 18, x- 1ex). : Murayama, 1929b: 43 ( - ); Nakayama and Okamoto, 1940: 198 (bicaloreus: ).

: : 169 Tribe Melolonthini Leach in Samouelle, 1819: 189. Melolonthidæ Leach in Samouelle, 1819: 189.,. 10 4 7.. 2 3 1, 1. : Melolontha Fabricius, 1775: 31. : 16 ( 2 ). : 200 ( 3 ). :. : Murayama (1954), Kim (2001b). 1.. 7, 5 Polyphylla laticollis manchurica. 2 Melolontha 2 2. 10. M. incana 26. M. insulana Genus Polyphylla Harris, 1841: 30. Macronoxia Crotch, 1873., 7, 5. 2 3,. 2,. Lined June Beetles. 2 1

170.. : Melolontha variolosa Hentz, 1830 (Young, 1988 ). : 65 (Loebl and Smetana, 2006), 32 (Smith, 2002) ( 1 ). :. : Nijiima and Kinoshita (1923), Dewailly (1948). 81. Polyphylla laticollis manchurica Semenov, 1900: 308 ( 17-69, 15-69, 20-69) Polyphylla laticollis Lewis, Ent. Month. Mag., 23: 231, 1882: Niijima and Kinoshita, 1923: 68; Okamoto, 1924: 174; Kamijo, 1932: 21; Miwa and Chûjô, 1939: 68; Murayama, 1954: 133; Cho, 1956: 77 ( ); 1957: 300; 1963: 218 ( ); 1969: 673; Cho et al., 1968: 264; KZS, 1968: 137; Kim et al., 1975: 243; 1982a: 156; 1982b: 276; Kim, 1978: 343; Stebnicka, 1980: 207; Park and Kim, 1993: 113; Kim, 1993: 65; 1998c: 356; Lie et al., 1997: 53. Polyphylla laticollis mandschurica Semenov, Horae Soc. Ent. Ross., 38: 308, 1900 (Manchuria): Dewailly, 1948: 108. Polyphylla laticollis mandshurica: Reitter, 1902: 271. Polyphylla laticollis manchurica Semenov, Horae Soc. Ent. Ross., 38: 308, 1900: ESK and KSAE, 1994: 151; Kim, 2001b: 47. Polyphylla laticollis mandsurica: Miwa and Chûjô, 1939: 68; Cho, 1957: 300; KZS, 1968: 137. Polyphylla laticollis manchurica: ESK and KSAE, 1994: 151. Polyphylla chinensis manchurica: Miwa and Chûjô, 1939: 68; Stebnicka, 1980: 207. : (KZS, 1968 2)...,... 10 7, 1.5. 6,. 2, 3,.,. Ad:.,.,. LPMB= 2: 1: 4. BL: 30 37 mm,

: : 171 BW: 16 19 mm. : Zhang, 1984: 67 (P. laticollis). :,,. :,,,. : NK-PB,, PN, WH, SK-GS, (,,,, 2 ), CN, GB. JJ. (24ex. 49 ): [SWU] : 2 1 ( : vi.1999); 49 ( : 21.iv.2005). [KU] : 1ex ( : xi.1935). [EWU] : 12ex ( : v vii.1955 1969). [HMNH- NK] 8ex (, : 8.vii.1982). : 1970 (Kim, 1998c: 351).,. Genus Melolontha Fabrioius, 1775: 31. Hoplosternus Guérin, 1838. Apropyga Medvedev, 1951 ( : Melolontha aceris Faldermann, 1835). 20 mm 10 7 2 3 6... 1, 1., 4 2, 1 1. : Scarabaeus melolontha Linnaeus, 1758. Syst. Ent., 32: 31. : 140 (Loebl and Smetana, 2006) ( 2 ). :. : Nijiima and Kinoshita (1923), Murayama (1954).

172 82. Melolontha incana (Motschulsky), 1853: 46 ( 7-70, 15-70, 21-70) Oplosternus incana Motschulsky, Etud. Ent., II: 46, 1853. Melolontha incana: Arrow, 1913: 401; Niijima and Kinoshita, 63; Saito, 1928: 12; Maruta, 1929: 368; Kamijo, 1932: 21; Haku, 1935: 57; Masaki, 1936: 260; Mochizuki and Tsunekawa, 1937: 91; Miwa and Chûjô, 1939: 69; Cho: 300; 1969: 674; KZS, 1968: 137 ( ); KSPP, 1972: 207; Kim et al., 1972b: 197; 1975: 243; 1982a: 156; 1982b: 276; 1984b: 328; Gu, 1973: 163; Kim, 1978: 346; Shin, 1979: 144; Yoon et al., 1979: 150; 1981: 47; 1990b: 110; Nam and Kim, 1982: 129; Kim et al., 1989: 176; 1991b: 67; 1991e: 179; 1994b: 111; 1996b: 127; 1997: 234; 1998: 170; 1999: 129; 2001: 132; 2002: 120; 2003: 126; 2004: 116; 2005: 79; Park and Kim, 1993: 113; Park et al., 1993: 178; 137; Kim, 1993: 65; 1995a: 164; 1995b: 139; 1996c: 174; 1998c; 357; 2000: 132; 2001b: 48; ESK and KSAE, 1994: 150; Lie et al., 1997: 59. Hoplosternus incanus: Stebnicka, 1980: 261. : (Gu, 1973), (KSPP, 1972). :... 10 7 2, 6.,. 3 3., 1. Ad:.. 1.7. 2.. BL: 26 33 mm, BW: 13 16.5 mm. : Murayama, 1931: 48; Medvedev, 1952: 98 (H. incanus); Zhang, 1984: 59 (H. incanus). :,,. :,,,. : NK-HB,, JG, HN,, PB,, PY, WN, GW, SK- (15 ) 80 100 700. (458ex): [SWU] : 12ex (,,,, : vii viii.1980 2000); 5ex ( : 19.viii.2001); 2 ( : 20.vii.2009). : 43ex (,,,,,,,,,,,,,, : vii ix.1977 1998). : 13ex (,,,,, : viii x.1987 2000); 3ex ( : 27.ix.2000); 1ex ( : 5.viii. 2002). : 2ex ( : 29.vi.1998); 19ex (,,,,,,, : vi ix.1981

: : 173 1998). : 1ex ( : 20.viii.1989); 4ex (, : vi viii.1990 1996). [NAAS] NK-1ex ( : vii.1924). : 21ex (,,,,, : vii ix.1981 1994). : 61ex (,,,,, : vi ix.1922 1997). : 44ex (,,,,,,, : vii viii.1974 1998). : 74ex (,,,,,, : vii viii.1922 1997). : 5ex (,, : vii viii.1992, 1993). : 3ex (3.vi.1984). [SNU] : 3ex ( : viii.1996, 1997). : 20ex (,,,,, : v ix.1982 1997). : 1ex ( : 29.vi.1994). : 1ex ( : 25.viii.1996). : 5ex (, : vii viii.1968, 1970, 1994). [KU] : 2ex (, : vii.1975, 1981). : 22ex (,,,,, : v x.1929 1989). : 5ex (,, : 28.vii ix.1957 1988). : 2ex ( : vii.1967); 2ex ( : vii.1976). : 2ex (, : viii.1972 1982). [EWU] : 15ex (,,,,, : vii viii.1954 1994). : 35ex (,,,,,, : vi x.1954 1981). : 14ex (,,,, : vii, viii.1969 1971). : 11ex (,,,, : vii viii.1971 1983). : 6ex (,, : vi viii.1959 1974). [HSU] : 3ex ( : vii, viii.1994, 1997). : 384, 5 10.(v- 2, vi- 18, vii- 196, viii- 147, ix- 18, x- 3ex). : Saito, 1928: 12; Kim and Kang, 1993: 415. 83. Melolontha insulana (Moser), 1918: 246 ( 7-71, 15-71, 21-71) Hoplosternus insulana (Moser), Stett. Ent. Zeit., 79: 246, 1918. Melolontha insulana: Kim, 2001b: 49. :. ( ), 26 2.,..,,.,, 2. 3 1,..,. BL: 29 30 mm, BW: 14 15 mm ( 1 BL: 23.5 mm, BW: 11.5 mm). :,. :?. : [SWU] 1 (label ).

174 Melolontha (= ) 2 H. incana. H. incana, incana. 1. Melolontha frater Arrow, 1913 Melolontha frater Arrow, Ann. Mag. Nat. Hist.: 400, 1913: Nagaoka, 1938: 25; Murayama, 1954: 131; Cho, 1969: 674 ( ); Hyun and Woo, 1969: 193 ( ); KSPP, 1972: 207; Kim, 1978: 345; Yoon and Nam, 1979: 150; Kim and Nam, 1981: 125; 1984b: 328; 1985: 105; Nam and Kim, 1982: 129; Park and Cho, 1986: 128; Kim et al., 1991b: 179; 1991e: 67; Park and Kim, 1993: 113; Park et al., 1993: 178; 137; ESK and KSAE, 1994: 150; Lie et al., 1997: 56. 2. Melolontha japonica Burmeister, 1855 Melolontha japonica Burmeister, Handb. Ent., IV(2): 420, 1855: Cho et al., 1967: 197; Kim, 1956: 9; 1958: 98; 1960: 27; Gu, 1973: 73 ( ); Kim and Nam, 1984b: 328 ( ); Park and Kim, 1993: 113; ESK and KSAE, 1994: 150 ( ). Hoplosternus japonicus Harold, 1875: Bates, 1888: 373; Stebnicka, 1980: 207. (= ) 1. Cyphochilus farinosus Waterhouse, 1867 Cyphochilus farinosus Waterhouse, Ent. Month. Mag., IV: 143, 1867: Stebnicka, 1980: 207; ESK and KSAE, 1994: 150.. 2. Pollaplonyx flavidus Waterhouse, 1875 Pollaplonyx flavidus Waterhouse, Trans. Ent. Soc., Lond.: 105, 1875 (Japan); Park et al., 1993: 178 (

: : 175 ),. Subfamily Sericinae Kirby, 1837: 128. Sericidae Kirby, 1837. (, Tibial Spurs) 2,,, 5,..,.,. Sericina 1 11 50. (Kim and Kim, 2003a, b) 4 5 27. : Serica MacLeay, 1819: 146. : Diphucephalini, Ablaberini (=Camentini), Sericini 3, Ablaberini, Sericini 2 (Sabatinelli and Pontuale, 1998) ( Sericini 1 ). Sericini : Phyllotocina, Trochalina, Sericina 3 (Sabatinelli et Pontuale, 1998) ( Sericina 1 ). : 200 (Ahrens, 2007a) ( 5 ). : 3,500 (Ahrens, 2007a) [ 725 (Sabatinelli and Pontuale, 1998)] ( 27 ). :. : Brenske (1897), Reitter (1902), Moser (1915, 1915b), Sawada (1937b), Murayama (1935, 1938b, 1941, 1941a), Medvedev (1952a), Kim and Lee (1991f, 1997b), Kim and Kim (2003a, 2003b, 2003c), Ahrens (2007, 2007a, 2007b); Gastroserica herzi=murayama (1930b); Maladera infuscata=ahrens (2003). 1.. 10 4, 4 3. gen. Gastroserica

176., 2,,. G. herzi 2 2. 3 4 3., ( ). 9, 3., gen. Paraserica.,,., P. grisea.,. 9 3 gen. Maladera 4.,. 9 4 3 gen. Sericania, 9 5 5., 2/3.., gen. Nipponoserica,.,. gen. Serica Genus Gastroserica Brenske, 1897b: 412. 10 4, 4 3,,,. 6. 1. : Serica marginalis Brenske, 1894 (Nomura, 1973 ). Gastroserica sulcata Brenske, 1898 ( ).

: : 177 :. : 6 ( 1 ). :. : Murayama (1930b), Kim (2001b). 84. Gastroserica herzi (Heyden), 1887: 264 ( 8-72, 15-72, 21-72) Serica herzi Heyden, Horae Soc. Entom. Ross., 21: 264, 1887: Bates, 1888: 372. Microserica hertzi: Reitter, 1902: 143. Gastroserica herzi: Brenske, 1897: 414; Niijima and Kinoshita, 1923: 36; Murayama, 1930b: 58; 1935: 4; 1937: 34; 1938: 15; 1954: 74; Mochizuki and Tsunekawa, 1937: 91; Nagaoka, 1940: 474; Miwa and Chûjô, 1939: 60; Cho, 1957: 297; 1969: 663 ( ); Hyun and Woo, 1969: 193; Nakane, 1973: 63; Kim et al., 1972a: 83; 1974a: 229; 1974b: 108; 1975: 239; 1976: 102; 1982a: 129; 1982b: 276; Kim, 1978: 338; Stebnicka, 1980: 250; Kim, 1981: 344; 1992: 105; 1993: 65; 1995a: 164; 2001b: 55; Yoon and Nam, 1990b: 110; Kim et al., 1986: 104; 1987b: 505; 1991b: 67; 1991e: 179; 1994b: 111; 1996: 58; 1996a: 127; 1999: 129; 2002: 120; 2003: 126; 2004: 116; Park and Park, 1987: 673; Park and Kim, 1993: 112; Park et al., 1993: 178; Lee et al., 1994: 147; ESK and KSAE, 1994: 151. : (Cho, 1969 10). :.,, 2,,..,.,. 10 4,.,...,., 2,.. BL: 6 8.5 mm, BW: 3.5 4.5 mm. : Kim and Kim (2003c). :,.

178 :,,. : NK-HB, HN,,, PB, PN, GW, SK- (5 ) 90 130 2,000. (1,604ex): [SWU] : 75ex (,,,,,,,, : vi viii. 1976 1996); 2ex ( : 20.vi.2001); 2ex ( : 10.vii.2009). : 358ex (,,,,,,,,,,, : v ix.1958 1999). : 2ex [ (,, ): 5.viii.2002]; 17ex (17.vi.2003); 40ex (,,,,,,, : v viii.1989 1999). : 46ex ( : 29.vi 3.vii.1998); 56ex (,,,,,,,,,,,, : v viii.1965 1998); 1ex ( : 6.vi.2008). : 78ex (,,,,,,, : v viii.1985 1994). [NAAS] : 6ex ( : vi viii. 1980 1989). : 3ex (,, : v vii.1980 1983). : 10ex ( : 10.vi.1975). : 5ex (, : vi.1973, 1974). [SNU] : 1ex ( : 17.vii.1962); 1ex ( : vi.1992). : 188ex (,,,,, : v, vi.1982 1997). : 5ex ( : vi.1992); 1ex ( : vi.1973); 2ex ( : 3.vii.1983). : 1ex ( : 10.vii.?). : 1ex ( : vi.1991). : 122ex (,,,,,, : vi viii.1968 1997). [KU] : 16ex (,,,,,, : iv vii.1968 1994). : 341ex (,,,,,,,, : iv viii.1935 2000). : 13ex (,,, : vi vii.1960 2000). : 110ex (,,,,,,,, : iv viii. 1933 1996). : 216ex (,,,,,,, : v viii.1957 1998). [EWU] : 2ex ( : vi.1973, 1975); 2ex ( : vii.1983). : 28ex (,,,, : v vii, ix.1966 1989). [HSU] : 23ex ( : v vi.1994 1997). [HMNH-NK] : 1ex ( : 14.vii.1982). : 4ex ( : 31.vii.1982). : 1ex ( : 29.v.1970); 2ex (4.viii.1975); 3ex (9.vii.1977); 17ex (17 22.vi.1988); 1ex (11.vii.1991).?: 1ex (Soktan: 5.vii.1956); 2ex (Tschon-Bon- San: 1.vii.1956). : 1,402, 4 9. (iv- 11, v- 107, vi- 725, vii- 477, viii- 78, ix- 4ex). : Kim and Kang, 1993: 415. Genus Paraserica Reitter 1896: 183.,., ( ). 9, 3.,. P. taiwana 2. S. polita grisea. : Serica grisea Motschulsky, 1866: 171 ( ).

: : 179 : 2 ( 1 ). : (, ). 85. Paraserica grisea (Motschulsky), 1866: 171 Serica grisea Motschulsky, Bull. Soc. Nat. Hist. Moscou, 39, I: 171, 1866: Sawada, 1937b: 12. Paraserica grisea: Miwa and Chûjô, 1939: 55; Cho, 1957: 296; KZS, 1968: 136 (geisea: ); Stebnicka, 1980: 206; Yoon and Nam, 1986: 159; Kim, 2001b: 60. : ( Serica polita).,.,,,,,.. 9 3, 2.,. ( ).,. (Serica polita). BL: 7 9 mm, BW: 4 5 mm. : Gokan (1982). :,. :?. : NK-HB (, 1, 19.vi.1935);? (Sanbo, 1, 8.vi.1936, coll. Takeuchi): Sawada, 1937; (Yoon and Nam, 1985: ). :. :. Yoon (1985), Murayama (1954) Sawada Serica polita. Genus Maladera Mulsant and Rey, 1871: 599. Neoserica Brenske, 1894. Autoserica Brenske, 1897. Lepiserica Brenske, 1901.

180 Microserica Medvedev, 1952. Eumaladera Nomura, 1967.,. 10, 1, 1.5 9,.,. 3.. Autoserica, Aserica, Serica, Maladera, 30 14. : Scarabaeus holosericea Scopoli, 1772. : 500 ( 170 )( 14 ). :,,. : Brenske (1897), Reitter (1902), Moser (1915, 1915b), Sawada (1937b), Murayama (1954), Kim and Lee (1991f, 1997b), Kim (2001b), Kim and Kim (2003a, 2003b, 2003c), Ahrens (2007, 2007a, 2007b). 1.,. coreana,. 2 2., 3. 6 3. 8 mm. 4 7.5 mm. 5 4.. verticalis. castanea koreana 5.. ovatula,. aureola

: : 181 6. 1 7 9 7., renardi. 8 8. 2., ( ) 1 holosericea,., 1 schoenfeldti 9. 3 4 gibbiventris 10 10.,.. infuscata,. 11 11. fusania 12 12., 10 opaciventris, 9 10 13 13. 8 mm, 1 orientalis 8 mm, 14 14., 1 cariniceps, laboriosa 86. Maladera holosericea (Scopoli), 1772: 77 ( 8-73, 15-73, 21-73) Scarabaeus holosericea Scopoli, Ann. Hist. Nat. V: 77, 1772. Serica holosericea: Murayama, 1935a: 2; 1937a: 33; 1938b: 10; 1954: 52; Cho, 1969: 656 ( ). Maladera holosericea: KZS, 1968: 136; Kim and Lee, 1997b: 122; Kim, 2001b: 69. Maladera (Maladera) holosericea: Stebnicka, 1980: 225. Serica holoserica ( ): Eguti, 1932: 57; Kim and Kim, 1974b: 229; Kim and Nam, 1982a: 154. Maladera holoserica ( ): Miwa and Chûjô, 1939: 54; Cho, 1957: 124; Kim et al., 1987c: 505; 1991b: 67; 1996a: 127; Kim, 1992: 105; 1993: 65. : (KZS, 1968; Kim and Nam, 1982a).

182 (M. schoenfeldti),..,.. 10 2,.,.,,. 1., (, Spurs) 1. Ad:,,,.,. BL: 7 8.7 mm, BW: 4 5 mm. : Kim and Kim (2003c). : Medvedev, 1952: 148. :,,,,. :,,,. : NK-HB, HN,,, PB,,, PY, WH,,,, GW,, GG, SK- (7 ) 30 40 200. (149ex): [SWU] : 3ex (,, : v viii.1984 1999); 1ex ( : 4.v.2003). : 39ex (,,,, : iv viii.1966 1999); 2ex ( : 26.v, 29.ix.2003). : 1ex ( : v.1987). : 1ex ( : iv.1983). : 1ex ( : v.1992); 1ex ( : 30.vi.2003). : 2ex (, : v.1976, 1993); 1ex ( : 1.v.2009). [NAAS] : 25ex ( : iii viii.1921 1989). : 3ex (, : v vii.1975 1984). [SNU] : 1ex ( : 20.v.1983). : 22ex ( - - : iv vi.1981 1990). : 1ex ( : 19.v.1990). : 1ex ( : 12.v.1990). : 1ex ( : 21.vii.1981). [KU] : 3ex (, : iv, v.1987 1990); 1ex ( : v.1966). : 1ex ( : vi.1996). : 7ex ( : v.1988); 1ex ( : v.1991). [HSU] : 2ex ( : v, vi.1994, 1997). [EWU] : 1ex ( : 14.viii.1980). : 10ex (,, : v viii.1964 1993). [HMNH-NK] : 1ex ( cemetery: 8.x.1978). : 6ex ( : 21, 22.v.1985); 6ex (23 28.v. 1991). : 1ex (Kyollyong reservoir: 30.ix.1978). : 1ex ( : 29.v.1970); 1ex (23.x.1987); 1ex (12.vi.1991). : 108, 3 8, 10, 11. (iii- 8, iv- 9, v- 49, vi- 30, vii- 2, viii- 7ex, x- 2, xi- 1ex). 87. Maladera coreana Kim and Kim, 2003: 83 Maladera coreana Kim et Kim, Ins. Koreana, 20(1): 83, 2003. Eumaladera coreana: Loebl and Smetana, 2006: 230.

: : 183 Maladera nitidiceps Nomura, 1967 ( ): Stebnicka, 1980: 258 260. :.,.,. 10,..,.,.,.,,. :,. Ad:,.,.. BL: 8 9 mm, BW: 4.5 5.3 mm. : Kim and Kim (2003c). :. :?,,. :. (19ex, ): [SWU] SE: 1 ( : 8.vi.1989); 1 ( : 30.iv.1998); 1 1 ( : 11, 20.vi.2001); 1 ( : 21.vi.1999); 1 ( : 14.iv.1996); 1 ( : 19.v.1988); 1 ( : 1.viii.?); 1 ( : 1.ix.1993). : 1 ( : 24.vi.1986); 1 ( : 13.vi.2000); 1 ( : 20.v.1970). : 1 ( : 7.vi.1997); 1 (29.vii.1979). : 1 ( : 23.vii.1990). : 3 (, : 8 9.viii.1994). : 1 ( : 23.viii.1984). : M. nitidiceps Nomura Stebnicka (1980)... 88. Maladera verticalis (Fairmaire), 1888: 118 ( 8-74, 15-74, 21-74) Serica verticalis Fairmaire, Rev. d Ent., vii: 118, 1888: Murayama, 1954: 59; Cho, 1969: 659 ( ); Kim et al., 1974b: 107; 1982: 155; Kim, 1981: 344. Aserica verticalis: Murayama, 1935a: 3; 1938a: 14; 1941: 20; Cho, 1957: 125; KZS, 1968: 136. Maladera (Maladera) verticalis: Stebnicka, 1980: 207. Maladera verticalis: Kim et al., 1991a: 192; 1991b: 67; 1991e: 179; 1992b: 153; 1997b: 127; 1999: 129; 2001: 132; 2002: 120; 2003: 126; 2004: 117; Kim, 1992: 105; 1993: 65; 1996: 58; 1996a: 127; 1996c: 174; 1998: 170; 2000: 132; 2001b: 66; ESK and KSAE, 1994: 151.

184 : (KZS, 1968; Kim et al., 1974), (Kim and Nam, 1982). :.,..,., 10. 1. 1.. 2,, 1/4. 1 (, upper spur), 2.. Ad:,. BL: 8 9.5 mm, BW: 4.3 6 mm. : Kim and Kim (2003c). : Zhang, 1984: 92. :,,. :,,,. : NK-HB, HN,, PN, PY, GW, WH, ; GG,? Eishû, Kyôjô, Naizôzan, Kanyó, SK- 90 120 1,000. (943ex): [SWU] : 98ex (,,,,,, : vii viii.1974 2000); 5ex ( : 19.viii.2001). : 66ex (,,,,,,,, : v xi.1972 2000). : 46ex (,,,,,,,,, : vi viii.1964 2000); 3ex (, : 27.ix.2000); 15ex (5, 28.viii.2002). : 95ex (, : 29.vi 3.vii.1998); 83ex (,,,,,,,,,,,,,,,,,, : iv viii.1969 2000). : 32ex (,,,,,,,,,, : vii viii.1969 1998). : 3ex (vii viii.1985 1996). [NAAS] : 1ex ( : 7.x.1974). : 1ex ( : 1976); 266ex ( : iii viii.1973 1989). : 50ex (,,, : vii viii.1975 1988). [SNU] : 11ex ( : iv vi, ix.1964, 1989 1993); 1ex ( : 15.ix.1990). : 5ex ( : vii, xi.1983). : 1ex ( : 30.vii.1994). : 27ex ( : vii, viii,1968 1981); 1ex (24.v.1994); 6ex (, : vi.1994).?: 3ex. [KU] : 3ex (Mt. : v.1918); 3ex (, : vii, viii.1958, 1992). : 2ex ( : vii, viii.1975, 1989). : 3ex (, : vii viii.1957 1994). : 7ex ( : vii.1988). : 1ex ( : viii.1960). : 2ex ( : vi.1986). : 3ex (vii.1966 1973).?: 4ex (Shoko: vi.1914); 1ex (Jujibong: vii.1988). [HSU] : 6ex ( : v vi.1996, 1997). [EWU] : 3ex ( : viii.1979, 1981); 1ex ( : 17.vii. 1975). : 17ex [,,, (7 ): vi ix.1954 1993]. : 1ex ( : 29.vii.1983). : 1ex ( : 23.vii.1982). : 1ex (13.vii.1979). [HMNH-NK] : 7ex ( : 13 18.vii.1982); 2ex (17.viii.1989). : 33ex ( : 16 29.vii.1975); 1ex (25.ix.1978); 1ex (8.vii.1982); 5ex (28.vii.1982); 2ex (29, 30.vi.1991); 1ex ( : 10.vii.1982). : 3/5ex ( : 1 11.vii./1 15.viii.1956). :

: : 185 1ex ( : 31.vii.1982). : 4ex ( : 4 8.viii.1975); 4ex (22.vii.1982). : 2ex ( : 30.vii.1982); 2ex (25.viii.1989).?: 4ex (Tschondtsin: 29.vii.1956). : 796, 3 10. (iii- 2, iv- 1, v- 25, vi- 64, vii- 495, viii- 144, ix- 14, x- 1ex). 89. Maladera castanea koreana Kim and Kim, 2003: 90 ( 8-75, 15-75, 21-75) Maladera castanea koreana Kim and Kim, Ins. Koreana, 20(1): 90. Autoserica castanea Arrow, Ann. Mag. Nat. Hist., 8(12): 398, 1913: Cho et al., 1967: 198. Aserica castanea: Eguti, 1932: 58; Murayama, 1935a: 3; 1937a: 33; 1938b: 14; Miwa and Chûjô, 1939: 55; Cho, 1947: 65; 1957: 124. Serica castanea: Murayama, 1954: 44; Cho, 1969: 654 [(sic): castanae, ]. Maladera castanea: Nomura, 1969: 79; Shin and Joo, 1977: 88; Stebnicka, 1980: 253; Nam and Kim, 1982: 129; Kim, 1983: 83; 2001: 75; Kim et al., 1984: 169; 1991b: 67; 1997b: 127; 1998: 170; 2004: 117; Kim et al., 1985: 105; Park and Kim, 1993: 112; Park et al., 1993: 178. : castanea-,. castanea koreana-... 10,.,.,.,. ( ) 1. 1 2, 1.,., 1. Ad:.,. BL: 9 10 mm, BW: 5 6 mm. : Kim and Kim (2003c). : -,,,,,,. : -,. : M. castanea - 2 ( : 17.iv, 6.ix.1929), 1 ( : 12.viii.1937) 30 ( ). M.c. koreana-. (53ex): [SWU] : 1 [ : 26.vii.1982 ( )]. : 2 ( : 2, 3.viii.1996); 1 ( : 10.vii.1998). : 1 ( : 14.vi.1990); 1 ( : 16.ix.1990); 1 ( : 5.viii. 1996); 1 ( : 13.viii.1992); 2 (, : 13.vii.1975, 13.vi.1999); 1 ( : 5.vii.1986); 1 (29.vii.1993); 1 ( : 13.viii.1976). : 1 ( : 21.vii.1981); 3 ( : 6.viii.1990). :

186 1 (, : 22.vii.1994); 1 ( : 25.vii.1994); 2 ( : 28, 29.vii.1979); 1 ( : 19.vii.1999). : 1 ( : 20.vi.1992); 2 ( : 5.viii.1998); 1 (, : 22.vii.1997); 1 ( : 1.vii.1998); 1 ( : 10.vii.1977). : 1 ( : 15.viii.1987); 2 ( : 6, 27.vii.1984); 2 ( : 30, 31.vii.1981); 2 ( : 22, 24.vii.1985); 1 ( : 16.viii.1987); 4 ( : 15 18.viii.1985); 1 ( : 5.vii.1984). : 1 ( : 1.viii.1974); 1 ( : 28.vii.1988). : 1 ( : 19.viii.1982); 2 (1 29.viii.1998); 1 ( : 10.viii.1993)- ; : 1ex ( : 24.vii.2000). : 1ex ( : 4.vii.2003). : 2ex ( : 16 18.vi.2003). : 1ex ( : 5.vi.2008). : 1ex (26.vii.2003). : (M. verticalis),. 90. Maladera ovatula (Fairmaire), 1891: 195 ( 8-76, 15-76, 21-76) Serica ovatula Fairmaire, Ann. Soc. Ent. Belg., 35, Compt. Rend.: 195, 1891: Murayama, 1954: 64; Cho, 1969: 660; Kim and Kim, 1974b: 108; Kim and Nam, 1982a: 155. Autoserica ovatura: Niijima and Kinoshita, 1927: 5. Aserica ovatula: Murayama, 1938a: 14; Miwa and Chûjô, 1939: 57; Cho, 1957: 297; KZS, 1968: 136; Park and Han, 1992: 138. Maladera (Maladera) ovatula: Stebnicka, 1980: 207. Maladera ovatula: Kim, 1981: 344; 1992: 104; 1993: 65; 2001b: 67 ( ); Yoon and Nam, 1989: 140; Kim et al., 1989: 176; 1991b: 67; 1994b: 111; 1996: 58; 1997b: 128; 2001: 132; 2002: 120; 2004: 117; ESK and KSAE, 1994: 151. : (KZS, 1968). :., (Serica brunnea),.,. 10 1.4,.,...,,., 1,.,,.. Ad:

: : 187. ( ),,. BL: 7 8 mm, BW: 4 5 mm. : Kim and Kim (2003c). : Zhang, 1984: 92. :,,,. : (?),,,. : SK-GS,,, JB, JN,,,? Shinseimen. (161ex): [SWU] : 1ex ( : 24.vii.2000); 2ex ( : vi, vii.1981, 1996); 2ex ( : 19.viii.2001). : 19ex (,,,,,, : vi viii.1972 1998). : 3ex (, : 24.viii, 27.ix.2000). : 5ex (, : vi, vii.1979, 1994). : 1ex ( : 5.vi.2008); 1ex ( : 1.ix.2003). : 2ex (, : vii, viii.1974, 1996). : 1ex ( :?). : 1ex (26.vii.2003). [NAAS] : 1ex ( : 23.vi.1977). : 2ex ( : 19.viii.1976); 83ex ( : vii viii.1972 1976). : 2ex (, : vii.1975). : 1ex (22.vi.1975). [SNU] : 4ex (, : viii, ix.1964, 1993, 1996). : 1ex ( : 20.vii.1981); 1ex ( : 15.vii.1968). [KU] 1ex ( : iv.1991); 1ex ( : ix.1964). [EWU] : 1ex ( : 14.viii.1980); 1ex ( : 23.vii.1971). : 12ex [,, (7 ): vi x.1966 1975]. : 1ex ( : 6.vi.1969). [HMNH-NK] 6ex ( : 16.vii.1975). : 2ex ( : 20 25.vii.1956). : 2ex ( : 4.viii.1975). : 133, 4 9. (iv- 1, v- 1, vi- 7, vii- 69, viii- 49, ix- 6ex). 91. Maladera aureola (Murayama), 1938: 19 ( 8-77, 15-77) Aserica aureola Murayama, Annot. Zool. Jap., 17(1): 19, 1938: Murayama, 1938a: 19. Serica aureola: Murayama, 1954: 64; Cho, 1969: 661 ( ); Kim and Nam, 1982: 155. Maladera (Maladera) aureola: Stebnicka, 1980: 207. Maladera aureola: Kim et al., 1991b: 67; 1997b: 128; Kim, 2001b: 65. : (, ).,,..,. 10,.. 1,., 2, 1/4 1. 1 ( ), 2.

188 Ad:,. S. BL: 6.7 9 mm, BW: 3.9 5.6 mm. : Kim and Kim (2003c). :,. :,. : SK-GW, GS,,,, CN Chôchiin (= ), GB Eishû (=? ), Kôrei(= ), GN,, JB Seichû (=? ), JN. (17ex): [SWU] : 1 ( : 27.vi.1984). : 1 ( : 1.vi.1987). : 1 ( : 16.viii.1984); 1 ( : 20.vii.1984). [NAAS] : 11ex ( : v viii.1927 1984). [SNU] : 1ex ( : 15.vii.1976). 92. Maladera renardi (Ballion), 1870: 339 ( 8-78, 15-78, 21-78) Serica renardi Ballion, Bull. Soc. Nat. Hist. Moscou., XLIII: 339, 1870: Murayama, 1935b: 2; 1937a: 33; 1938b: 11; 1954: 54; Sawada, 1937a: 9; Cho, 1969: 657; Kim and Kim, 1972a: 84; Kim and Nam, 1982a: 154. Maladera renardi: Miwa and Chûjô, 1939: 55; Cho, 1957: 124; Nomura, 1967: 53; 1973: 128; KZS, 1968: 136 ( ); Stebnicka, 1980: 254; Kim, 1983: 83; 1992: 105; 1993: 65; 2001b: 68; Kim et al., 1991b: 67; 1994b: 111; 1997b: 123; 1998: 170; 2003b: 84; Bae and Moon, 1993: 147. Serica motschulskyi Brenske, Berl. Ent. Zeit., 52: 379, 1897 (Corea) (Syn.): Niijima and Kinoshita, 1923: 22; Murayama, 1937a: 33; 1938b: 11; 1954: 58; Sawada, 1937a: 9; Mochizuki and Tsunekawa, 1937: 93; Miwa and Chûjô, 1939: 53; Cho, 1957: 295; 1969: 658; Nomura, 1967: 52 (Syn. nov.); Kim and Kim, 1972a: 53; Kim and Nam, 1982a: 155. Maladera (Aserica) motschulskyi: Nomura, 1960: 58. Serica spissigrada Brenske, Berl. Ent. Zeit., 52: 430, 1897 (Japan) (Syn.): Niijima and Kinoshita, 1923: 23; Sawada, 1937a: 29; Murayama, 1938b: 12; 1954: 40; Nomura, 1967: 52 (Syn. nov.); KZS, 1968: 136; Cho, 1969: 652; KSPP, 1972; Kim et al., 1982a: 154. Maladera spissigrada: Miwa and Chûjô, 1939: 55; Cho, 1957: 296; Kim, 1981: 344. Maladera (Aserica) spissigrada: Nomura, 1960: 58. Serica nakayamai Murayama, Annot. Zool. Jap., 17(1): 16, 1938b (Syn.): Murayama, 1954: 57; Nomura, 1967: 52 (Syn. nov.); Cho, 1969: 658; Kim and Kim, 1972a: 83; Kim and Nam, 1982a: 155. : renardi= (Cho, 1969); motschulskyi= (Cho, 1969); spissigrada= (KZS, 1968), (KSPP, 1972); (Cho, 1969).

: : 189 :. (M. holosericea)..,. 10 1.5,..,., 1.3 1.5,. 1 ( ).,. Ad:,. 1. BL: 8 10 mm, BW: 6 mm. : Kim and Kim (2003c). : Medvedev, 1952: 150. :,,,,. :,,. : NK-HB,,, HN, YG, PB,, PY, WN, GW,,, SK-GW,,,, GS,,,,,,,,, CBN,,,, GB, JBN,,. (71ex): [SNU] : 14ex (,, - : 1985 1992); 1ex ( : ix.1934); 2ex ( : v, vi.1991, 1992); 1ex ( : 23.iv.2005). [KU] : 1ex ( : v.1966). : 1ex ( : v.1992); 1ex ( : vi.1961); 1ex ( : vi.1974). : 1ex ( : vi.1991). : 1ex ( : vi.1997); 3ex ( : vi.1996). : 1ex ( : v.1991). [HSU] : 28ex ( : iv vi.1994 1997). [EWU] : 1ex ( : 26.v.1962). : 13ex [,,,, (4 ), : iv vi.1956 1989]; : 1ex ( : 20.v.1988). : Saito, 1928: 13; Murayama, 1938a: 259. 93. Maladera schoenfeldti (Murayama), 1937: 33 ( 8-79, 15-79, 21-79) Serica schoenfeldti Murayama, J. Chosen Nat. Hist. Soc., 22: 33, 1937: Murayama, 1938b: 11; 1954: 56; Miwa and Chûjô, 1939: 54; Cho, 1957: 295; 1969: 657; Kim and Nam, 1982a: 154. Maladera (Maladera) schoenfeldti: Stebnicka, 1980: 258; Loebl and Smetana, 2006: 236. Maladera schoenfeldti: Kim et al., 1991b: 67; 1997b: 123; 2003b: 87; ESK and KSAE, 1994: 151 ( ); Kim, 2001b: 70.

190 : (KZS, 1968), (Kim and Lee, 1991b), (Cho, 1969). :.,.,,. 10,. 1.4,.,.., 2,. 1 ( ). Ad: (M. renardi).,. BL: 8 9.5 mm, BW: 4.3 5.4 mm. : Kim and Kim (2003c). :. :,,,. :,,, ( ), NK-,, SK-GW,,, GS,,,,,,,,,, CBN,,,,, GBN,,,,,,,,,,,,,,,, JBN,,, JJ,? Hoban ( ), Kankó, Kosei. (88ex): [SWU] : 1ex ( : vi.1974). : 19ex (,, : x, iv ix.1974 1998). : 7ex (,,,, : iv vii.1979 1999). : 20ex (,,,,,,,,,,,,, : iv ix.1977 1997). : 1ex ( : viii.1989). [SNU] : 4ex ( : iv vi.1986, 1990); 1ex ( : 17.v.1992); 1ex ( : 23.vi.1992). : 5ex (30.iv.1985).?: 1ex (Hoban [ ]: 26.vii.1936). [KU] : 6ex ( : vi.1996 1999); 1ex ( : vi.1994); 2ex ( : vi.1992). : 1ex ( : v.1968); 1ex ( : v.2000); 1ex ( : ix.1972); 1ex (Mt. Suraksan: v.1999). : 1ex ( : vi.1996); 1ex ( : vi.1990). : 3ex ( : vi.1972). [EWU] : 7ex (,, : v, vi.1974 1985). : 1ex ( : 8.v.1979). 94. Maladera gibbiventris (Brenske), 1897: 396 ( 8-80, 15-80, 21-80) Autoserica gibbiventris Brenske, Berl. Ent. Zeit. XLII; 396, 1897: Niijima and Kinoshita, 1927: 9; Murayama, 1935a: 3. Aserca gibbiventris: Murayama, 1938b: 12; Miwa and Chûjô, 1939: 56; Cho, 1957: 296; KZS, 1968: 136 ( ).

: : 191 Serca gibbiventris: Murayama, 1954: 36; Cho, 1969: 652; Kim and Kim, 1974b: 107; Kim and Nam, 1982a: 154. Maladera (Maladera) gibbiventris: Stebnicka, 1980: 254. Maladera gibbiventris: Kim et al., 1991b: 67; 1994b: 111; 1997b: 126; 1999: 129; 2001: 132; 2003: 126; 2003b: 87; Kim, 1992: 105; 1993: 65; 2001b: 74; ESK and KSAE, 1994: 151. :.,.,,.. 10 1.7, 1.1.. 3 4. Ad:,.. BL: 9 12 mm, BW: 5.5 7.3 mm. : Kim and Kim (2003c). :,,. :,,. : NK-HB, PB, Seishin (= ), PY,,,,, PN, WN, GW, SK- 70 90 200. (146ex): [SWU] : 2ex ( : vi.1984); 4ex (,, : v vi.1974 1992). : 36ex (,,,, : iv ix.1972 1998). : 4ex (,,, : v ix.1984 1999), 1ex (, : 18.v.2001); 2ex (5.viii.2002). : 1ex ( : 1.vii.1998); 16ex (,,,,,,,,, : 1983 1992); 1ex ( : 9.v.2003). : 12ex (,,,,, : v vii.1977 1998). [NAAS] : 1ex ( : 30.v.1974). : 2ex ( : viii.1972, 1973). [SNU] : 1ex ( : 5.v.1987); 3ex ( : iv vi.?, 1981, 1992). : 1ex ( : 24.vi.1995). [KU] : 11ex (,,,,, : iv x.1961 2000). : 2ex ( : vi.1997). : 4ex (,, : v vi.1988 1998). [HSU] : 2ex ( : v.1994, 1997). [EWU] : 1ex ( : 28.v.1977); 2ex ( : vi.1979). : 21ex [,,,, (9 ), : v x.1957 1990]. : 1ex ( : 5.vii.1978). [HMNH-NK] : 3ex ( : 17.v.1985); 4ex (23 28.v.1991). : 2ex ( : 17.v.1985); 1ex (31.v.1985). : 1ex ( : 24.v.1985). : 2ex ( : 6.viii.1975); 1ex (26.v.1985); 2ex (11.vii.1991). : 111, 4 10. (iv- 5, v- 61, vi- 30, vii- 10, viii- 5, ix- 3, x- 2ex). : Maruta, 1929: 367; Nakayama and Okamoto, 1940: 198.

192 95. Maladera infuscata (Moser), 1915: 34 ( 8-81, 15-81, 21-81) Autoserica infuscata Moser, Deut. Ent. Zeit.: 340, 1915: Niijima and Kinoshita, 1923: 29; 1927: 6. Aserica infuscata: Murayama, 1938b: 13; Miwa and Chûjô, 1939: 56; Cho, 1957: 297; KZS, 1968: 136 ( ). Serica infuscata: Murayama, 1954: 41; Cho, 1969: 653; Kim and Nam, 1982a: 154. Maladera (Maladera) infuscata: Stebnika, 1980: 207. Maladera infuscata: ESK and KSAE, 1994: 151; Kim et al., 1997b: 122; Kim, 2001b: 74; Loebl and Smetana, 2006: 237. Autoserica koreana Moser, Stett. Ent. Zeit., 53: 330, 1919 (, Chemulpo): Niijima and Kinoshita, 1923: 29; Ahrens, 2003: 3 (syn. nov.). Aserica koreana: Murayama, 1938b: 15; Miwa and Chûjô, 1939: 57; Cho, 1957: 297; KZS, 1968: 136 ( ). Serica koreana: Murayama, 1954: 65; Cho et al., 1968: 661. Maladera koreana: Stebnika, 1980: 207. Aserica okamotoi Murayama, Annot. Zool. Jap., 17 (1): 18, 1938b ( 5 st. 28 ex)(syn.): 1954: 35./ Ahrens, 2003: 3 (syn. nov.: Maladera). Serica okamotoi (Syn.): Cho, 1969: 651 ( ); Kim and Kim, 1974a: 229. Maladera okamotoi (Syn.): Stebnika, 1980: 258 (, 5ex: -.); Kim et al., 1991b: 67; 1996: 58; 1997b: 126; Kim, 1992: 105; 1993: 65; 2001: 76; ESK and KSAE, 1994: 151; Ahrens, 2003: 3 (syn. nov.). : koreana= (KZS, 1968; Cho et al., 1968), okamotoi= (Cho, 1969 8). :,.,,,.,. 10,. 1,.,.,, 2. 1 ( ),.. 1. Ad:,,.,.,. BL: 8 10.5 mm, BW: 5 6.5 mm.

: : 193 :, ( ),. :,,. : infuscata=sk-,,, ; okamotoi=nk-,,, Mt. Tshonbonsan, SK- 50 150. (121ex): [SWU] : 2ex (, : v vii.1983 1995); 4ex ( : vi.1984). : 10ex (,,, : v vii.1974 1995); 1ex ( : 26.v.2003). : 1ex ( : v.1987). : 5ex (,, : v ix.1983 1994). : 12ex (,,,,,,, : iv vii. 1974 1996). : 8ex (,, : v vii.1974 1994). [NAAS] : 2ex ( : 30.v.1974). : 4ex ( : v vii.1975 1984). [SNU] : 1ex ( : 6.vi.1987); 1ex ( : 18.iv.1992). : 1ex ( : 10.v.1992). : 1ex ( : vi.?). [KU] : 4ex ( : v.1968); 1ex ( : v.1986). : 6ex (,, : v vi.1959 1991). : 3ex ( : vi.1996). : 12ex (,, : v vi.1986 1998). [HSU] : 17ex ( : v vii.1994, 1997). [EWU] : 1ex ( : 14.viii.1980). : 1ex ( : 9.v.1965); 1ex ( : 26.v.1990); 1ex ( : 5.vi.1983); 3ex ( : v, vi.1972 1983); 1ex ( : 3.vi.1973); 1ex ( : 30.v.1983). : 5ex ( : v.1966, 1977). [HMNH- NK] : 2ex ( : 24 28.v.1991). : 2ex ( : 9, 14.viii.1971); 1ex (16.vii.1975); 1ex (19.ix. 1979). : 1ex ( : 1.vi.1970); 1ex (4.viii.1975); 1ex (10.vi.1991).?: 1ex (Mt. Tshonbonsan: 3.ix. 1956). : 115, 4 9. (iv- 1, v vii- 108, viii- 4, ix- 2ex). 96. Maladera fusania (Murayama), 1934: 35 ( 8-82, 15-82, 21-82) Aserica fusania Murayama, J. Chosen Nat. Hist. Soc., 19: 35, 1934: Murayama, 1935a: 3; 1938b: 13; Miwa and Chûjô, 1939; Cho, 1957: 296; KZS, 1968: 136 ( ). Serica fusania: Murayama, 1954: 38; Cho, 1969: 652; Kim and Kim, 1972a: 84. Maladera (Maladera) fusania: Stebnicka, 1980: 255. Maladera fusania: Kim et al., 1991b: 67; 1991d: 192; 1997b: 125; 2001: 132; ESK and KSAE, 1994: 151; Kim, 2001b: 76. Maladera (Omalodera) fusania: Loebl and Smetana, 2006: 236. : ( ).,... 10.. Ad:.. BL: 7.5 9.5 mm, BW: 5 6 mm.

194 : Kim and Kim (2003c). :,. :,,,. : NK-HN,, PY,,, WN,, GG, SK- 70 100 1,000. (805ex): [SWU] : 1ex ( : vii.1993); 52ex (,, : iv vi.1977 1987). : 120ex (,,,,,,,,,,,, : iv x.1956 1999); 2ex ( : 26.v, 5.vii.2003); 1ex ( : 4.vii.2003); 1ex ( : 10.vi.2008); 2ex ( : 26.vi.2009). : 13ex (,,,,, : iii viii.1977 1999); 1ex ( : 18.v.2001). : 56ex (,,,,,,,,,,,,,, : iii viii.1969 1994); 2ex ( : 19.v.2003); 5ex ( : 13.vii.2003). : 21ex (,,,,,,,,, : v vii.1976 1999); 1ex ( : 23.iv.2009); 1ex ( : 1.v.2009). [NAAS] : 3ex ( : iii v.1974 1982). [SNU] : 177ex (,,,,,, : iv vi.1965 1995). : 3ex (, : vi, vii.1967, 1992). : 1ex ( : v.1990). : 10ex (,, : vi, vii.1991). : 2ex (v.1992, vi.1988). [KU] : 5ex [Seiyori (= ): v ix.1932 1934]; 37ex (,,,,,, : v vii. 1942 1998). : 2ex (, : vi.1996). : 11ex (,,, : iv vii.1972 1993). : 1ex (vii.1966). [HSU] : 153ex ( : iv vi.1994 1997). [EWU] : 73ex ( : v viii.1959 1990); 1ex ( : 20.vi.1975). : 55ex (,,,,,,,, :v viii.1965 1991). : 1ex ( : 24.vii.1964). : 1ex ( : 10.vi.1983). : 1ex ( : 10.viii.1966); 1ex ( : 20.vii.1981). : 2ex (v.1981, vii.1985). : 281, 3 10. (iii- 4, iv- 17, v- 103, vi- 95, vii- 56, viii- 4, ix- 6, x- 1ex). 97. Maladera opaciventris (Moser), 1915: 355 ( 18-83, 15-83, 21-83) Autoserica opaciventris Moser, Deut. Ent. Zeit.: 355, 1915; Niijima and Kinoshita, 1923: 29. Aserica opaciventris: Murayama, 1938a: 12; Miwa and Chûjô, 1939: 57; Cho, 1957: 297; KZS, 1968: 136 ( ). Serica opaciventris: Murayama, 1954: 26 (GB ); Cho, 1969: 649. / : Kim and Kim, 1972: 83; Kim and Nam, 1982: 154; Lee et al., 1994: 147. Maladera (Maladera) opaciventris: Stebnicka, 1980: 207. Maladera opaciventris: Kim and Yoo, 1987: 505 ( ); 1997: 129; Kim, 2001b: 65; Loebl and Smetana, 2006: 237. : ( ).

: : 195.,,. 10,.,.,... BL: 8 9 mm. :. :,. : (Moser: 1915), Eishú (= : Murayama, 1938a), ( ), ( ). (2ex): [SWU] SE: 1ex ( : 28.viii.2006). : 1ex ( : 24.vii.1999). : Stadinger Bang-Hass, Murayama (1954) 1. Ahrens (2007) Eumaladera (Nomura, 1963). 98. Maladera orientalis (Motschulsky), 1857: 33 ( 8-84, 16-84, 2-84) Serica orientalis Motschulsky, Etud. Ent., VI: 33, 1857: Kolbe, 1886: 192; Heyden, 1887: 251; Niijima and Kinoshita, 1923: 21; Okamoto, 1924: 172; Murayama, 1938a: 259; 1938b: 10; 1954: 48; Masaki, 1936: 261; Sawada, 1937a: 25; Mochizuki and Tsunekawa, 1937: 93; Miwa and Chûjô, 1939: 53; Cho, 1957: 295; 1963: 217; 1969: 655; Cho et al., 1967: 197 ( ); 1968: 264; Kim and Kim, 1972a: 84; 1972b: 196; 1974a: 229; 1974b: 107; Kim and Nam, 1982a: 154; 1984: 328. Aserica orientalis: Reitter, 1902: 145. Maladera orientalis: Nomura, 1960: 58; 1969: 79; Stebnicka, 1980: 257; Kim, 1981: 344; 1992: 105; 1993: 65; 2000: 132; 2001b: 71; Kim and Nam, 1982b: 276; Kim et al., 1987a: 104; 1987b: 505; 1991b: 67; 1991d: 192; 1991e: 179; 1994b: 111; 1997b: 124; 1999: 129; 2001: 132; 2004: 117; Park et al., 1993: 178. Maladera (Omalodera) orientalis: Lobl and Smetana, 2006: 236. Serica salebrosa Brenske, Berl. Ent. Zeit., 17: 428, 1897 (Japan)(Syn.): Masaki, 1936: 261; Murayama, 1938b: 10 (Syn. nov.). : (Cho, 1963; Cho et al., 1968).,..,.,

196. 9.,.,.. 1 ( ),.. Ad:,.,.. BL: 6.5 8.6 mm, BW: 4 5.2 mm. : Kim and Kim (2003c). : Murayama, 1931: 20; Medvedev, 1952: 150; Zhang, 1984: 91 (S. orientalis). :,,,,,. :,,,. : NK-HB,,,,, HN,,,,, PB, PY,, PN,,,,, WN,, GW, GG,, SK- 80 120 1,200. (954ex): [SWU] : 46ex (,,,,,, : iv vii. 1974 1995). : 242ex (,,,,,,,, : iv x.1966 1997); 1ex ( : 4.iv.2004). : 27ex (,,,,,,, : v viii.1974 1996); 7ex ( : 18.v.2001); 1ex (19.vi.2003). : 2ex ( : 29.vi.1998); 47ex (,,,,,,,,,, : v viii.1970 1998). : 16ex (,,,,,,, : iv vii.1977 1994); 1ex ( : 7.v.2006). : 3ex (v.1997). [NAAS] : 5ex (, : v vi.1973 1986). : 125ex ( : iii viii.1923 1990); 1ex ( : 12.v.1985). : 2ex (, : iv, vi.1973, 1984). : 1ex ( : 11.v.1976). : 6ex (, : v viii.1975 1984). : 8ex (vi.1922 1984). [SNU] : 2ex (, : v.1983, ix.1997). : 58ex (,,,,,,, - - : v vi.1962 1992). : 6ex ( : vi, vii, ix.1983 1985); 2ex (, : vi.1992). : 1ex ( : viii.1994). : 1ex ( : v.1991). : 1ex ( : vi.1991). : 2ex (v.1992, vii.1990).?: 3ex. [KU] : 5ex (,, : iv vi.1957 1992). : 2ex [Seiryori (= ): vi, ix.1932, 1933]; 95ex (,,,,,,,,, : iv ix.1956 1998). : 2ex (, : v, vi.1987, 2000). : 7ex (,, : iv vi.1989 1996). : 15ex (,,, : v vi.1956 1998). [HSU] : 53ex ( : iv vi.1994 1997). [EWU] : 1ex ( : 3.vi.1972). : 136ex [ (17 ) : iv vi.1957 1990]. : 1ex ( : 17.v.1963). : 2ex (v.1988, vii.1985). [HMNH-NK] : 1ex ( : 25.v.1991). : 1ex ( : 17.v.1985); 1ex ( : 23.v.1970). : 3ex ( : 1 11.vii.1956); 2ex (1 15.viii.1956). : 1ex ( : 28.v.1970). : 3ex ( : 7.vi.1970). : 658, 3 10. (iii- 17, iv- 89, v- 237, vi- 236, vii- 57, viii- 18, ix- 3, x- 1ex). : Murayama, 1936: 1 6; 1938: 259 264; Nakayama and Okamoto, 1940: 200.

: : 197 99. Maladera cariniceps (Moser), 1915: 341 ( 9-85, 16-85, 21-85) Autoserica cariniceps Moser, Deut. Ent. Zeit.: 341, 1915: Niijima and Kinoshita, 1923: 238; 1927: 7. Aserica cariniceps: Murayama, 1938b: 13; Miwa and Chûjô, 1939: 55; Cho, 1957: 124; KZS, 1968: 136. Serica cariniceps: Murayama, 1954: 41; Cho, 1969: 653 ( ); Kim and Nam, 1982a: 155. Maladera (Aserica) cariniceps: Normura, 1967: 52. Maladera (Maladera) cariniceps: Nomura, 1973: 133; Stebnicka, 1980: 255. Maladera cariniceps: Kim et al., 1991b: 67; 1997b: 124; Kim, 1992: 105; 1993: 65; 2001b: 72; ESK and KSAE, 1994: 151. Maladera (Omalodera) cariniceps: Lobl and Smetana, 2006: 236. Aserica fusania Murayama, J. Chosen Nat. Hist. Soc., 19: 35, 1934 (, ): Nomura, 1967: 52 (Syn. nov.). : (KZS 1968, 3). : ( ).,..,.,. 10 9..,.,,.. Ad:,,,. BL: 8 10.5mm, BW: 5 6.5 mm. : Kim and Kim (2003c). :,,. :,,,. : NK-HB, HN, PB, PY,, WN, GG, SK- 80 100 700. (577ex): [SWU] : 7ex (,,, : v viii.1977 1996). : 171ex (,,,,,,,,,,,,, : iii ix.1966 2000); 1ex ( : 2.xi.2006). : 23ex (,,,,,,,,, : v viii. 1977 2000); 1ex ( : 24.viii); 2ex (27.ix.2000). : 56ex (,,,,,,,,,,,,,,, : iv viii.1978 1999). : 23ex (,,,,,,, : v viii.1976 1999). : 1ex (vii.1985). [NAAS] : 43ex ( : iii viii.1972 1990); 1ex [Kasan (= ): 6.iv.1922]. : 25ex ( : 12.iv.1996). :

198 1ex ( : v.1984). : 2ex (v, vi.1975). [SNU] : 1ex ( : v.1983). : 60ex ( : v vii.1965 1994); 6ex (,,,, : iv ix.1986 1994). : 2ex (, : vi.1992). : 1ex ( : vi.1994). : 1ex (iv.1986).?: 5ex [KU] : 1ex ( : vi.1999); 1ex ( : viii.1961). : 1ex [Seiyori (= ): ix.1932]; 10ex (,,,,, : v viii.1954 2000). : 1ex ( : vi.2000). : 1ex ( : vi.1997). : 2ex (, : vi.1990, 1996). : 9ex (,, : v.1988 1991). [HSU] : 22ex ( : iv vi.1994 1997). [EWU] : 1ex ( : 28.v.1977). : 82ex [,,,, (47ex. ): iv vi.1961 1991); 1ex (ix.1968). [HMNH-NK] : 1ex ( : 2.vi.1991); 4ex ( : 13, 14.vii.1981); 1ex ( : 7.vii.1982); 1ex (30.v.1985). : 1ex ( : 23.v.1970); 1ex ( : 27.vii.1982). : 1ex ( : 16.x.1987). : 375, 3 11. (iii- 16, iv- 60, v- 152, vi- 80, vii- 33, viii- 21, ix- 14, xi- 2ex). : Murayama, 1931: 259 (Aserica). Maladera (= ) 1. Maladera formosae (Brenske), 1898 Autoserica formosae Brenske, Berl. Ent. Zeit.: 210, 1898 (Formosa=Taiwan): Niijima and Kinoshita, 1923: 27; Aserica: Murayama, 1937a: 34; KZS, 1968: 136 ( ) 3; Serica: Murayama, 1954: 62; ESK and KSAE, 1994; 151 ( ) 2; Maladera: Stebnicka, 1980: 254 2 Niijima, Stebnicka. 2. Maladera japonica (Motschulsky), 1860 Serica japonica Motschulsky, Etud. Ent. IX: 15, 1860: Kolbe, 1886: 192 ( ); Hyun and Woo, 1969: 194 ( ); Cho et al., 1968: 264 (, ); Kim and Kim, 1972a: 83 ( ) 10; Autoserica: Nomura, 1967: 131 2; Maladera: Miwa, 1931: 289; Eguchi, 1932: 50 ( ); Paik, 1935: 57 ( ); Masaki, 1936 (, ) 25 Murayama (1954), Nomura (1973).

: : 199 3. Maladera kamiyai (Sawada), 1937 Serica kamiyai Sawada, Nippon no Kochu, 1(1): 27, 1937: Kim and Chang (1987a, ) Nipponoserica opacicarina. 4. Maladera laboriosa (Brenske), 1897 Aserica laboriosa Brenske, Berl. Ent. Zeit., XLII: 339, 1897: Murayama, 1937: 34 ( ); 1938: 12; Miwa and Chûjô, 1939: 57; Cho, 1957: 297; KZS, 1968: 136 ( ); Serica: Murayama, 1954: 43; Cho, 1969: 654; Maladera: Stebnicka, 1980: 207 Murayama (1954) 1937. 5. Maladera nitidiceps Nomura, 1967 Maladera nitidiceps Nomura, Ent. Rev. Jap., XIX(2): 55, 1967: Stebnicka, 1980: 258 (, ) M. coreana. 6. Maladera secreta (Brenske), 1897 Autoserica secreta Brenske, Berl. Ent. Zeit.: 341, 1897: Okamoto, 1924: 173 ( ); Morita, 1936: 861 ( ); Aserica: Murayama, 1938: 12; Miwa and Chûjô, 1939: 57; Takahashi, 1941: 230; Serica: Cho, 1957: 218 ( ); 1969: 651; Cho et al., 1968: 264; KZS: 136; Murayama, 1954: 35; Maladera: KSPP, 1972: 207 Murayama (1954) Okamoto, Morita (1936). 7. Maladera stridula (Brenske), 1897 Aserica stridula Brenske, Berl. Ent. Zeit.: 401, 1897: Murayama, 1935: 3 ( ); 1938: 12; 1954: 43; Serica: Cho, 1969: 654 ( ); Maladera: Stebnicka, 1980: 207 Murayama (1954) (1935).

200 8. Maladera thibetana (Brenske), 1897 Serica thibetana Brenske, Berl. Ent. Zeit.: 401, 1897: Nijiima and Kinoshita, 1927: 2 ( ); Murayama, 1935a: 2 ( ); 1938: 9; 1954: 34; Sawada, 1937: 9; Miwa and Chûjô, 1939: 54; Cho, 1957: 295; 1969: 651 ( ); KZS, 1968: 136 ( ) Thibet, Murayama (1954), Niijima. Genus Sericania Motschulsky 1860: 256. Mesoserica Matsumura, 1911: 118 ( : Mesoserica sachalinensis Matsumura, 1911 )., 9 10 4, 3.. 2 ( ), 2 3. 1/2 2/3.. Ahrens (2007b) 3 S. fuscolineata, 2 sister group. Heyden (1887) 8 1, 3 4. : Sericania fuscolineata Motschulsky, 1860: 136. : 70 ( 4 ). :. : Murayama (1935, 1937, 1941, 1954), Kim (2001b), Ahrens (2007, 2007a, 2007b). 1... yamauchii 2 2.,., koryoensis. 3

: : 201 3.. 4. 4, latisulcata.,. 4 5 4.5 5 fuscolineata 100. Sericania fuscolineata Motschulsky, 1860: 136 ( 9-86, 16-86, 22-86) Sericania fuscolineata Motschulsky, Schrenck s Reisen Amurl. II(2): 136, 1860: Heyden, 1887: 251; Niijima and Kinoshita, 1927: 13; Murayama, 1935: 3; 1937: 35; 1938: 15; 1954: 68; Doi, 1938: 97; Sawada, 1938: 10; Miwa and Chûjô, 1939: 58; Cho, 1957: 297; 1969: 662 ( ); Cho et al., 1967: 197; Kim et al., 1971: 160; 1981: 125; 1982a: 155; 1984: 328; 1985: 105; Kim, 1978: 383; Stebnicka, 1980: 252; Kim, 1981: 344; 1992: 105; 1993: 65; 1995a: 164; 1996c: 174; 2001b: 58; Kim et al., 1991b: 67; 1991f: 49; 1996a: 127; 1998: 170; 2002: 120; Park et al., 1993: 178; Loebl and Smetana, 2006: 246. Serica fuscolineata: ESK and KSAE, 1994: 151. : (Cho et al., 1967)..,,,..,. 9 4 2. 5. 3., 1.,., 1.. 2. 1 ( ). Ad: 1.2 mm,., 30.. BL: 8 11 mm, BW: 4.5 5.5 mm. : Kim and Kim (2003c). :,,,. :,,. : NK-HB,,,, ( ),,,,,,, HN

202,,, PY,, WN, GW,,,,,?-,, SK- 80 90 600. (525ex): [SWU] : 76ex (,,,.,,,, : v viii.1958 2003); 6ex ( : 20.vi.2001). : 52ex (,,,,,,,,,,, : iv vii.1957 2001). : 5ex (,, : iv v.1991 1999). : 116ex (,,,,,,,,,,, : v viii.1974 1999). : 49ex (,,, : v vii.1972 1997). [NAAS] NK-1ex [Sharei (= ): 21.viii.1922]. : 1ex ( : 10.vi.1973). : 23ex ( : v vii.1975 1981). : 1ex ( : 16.v.1975). : 1ex ( : 10.vi.1975).?: 2ex (Renen hunei: 1.iii.1927). [SNU] : 6ex ( : v vii.1991 1995). : 1ex ( : vi.1992); 3ex ( : v, vi.1982, 1991); 1ex ( : vii.1995); 1ex ( : 23.vi.1992). : 1ex ( : 17.v.1992). : 10ex (, : v, vi.1993, 1997); 5ex (,, : 23 5.vi.1995).?: 6ex [KU] : 42ex (,,,,, : v vi.1979 1999). : 15ex (,, : iv vi.1981 1997). : 10ex (,, : v vi.1987 2000). : 5ex (,, : vi.1981 1996). : 12ex (,,, : v vii.1983 1998). [EWU] : 3ex (,, : v, vi.1990 1995). : 32ex [,,,,, (6 ): v vi, ix.1959 1995]. : 1ex ( : 21.vi.1987). : 1ex ( : 17.v.1963). : 1ex ( : 6.v.1977). [HMNH-NK] : 1ex ( : 2.vi.1991). : 5ex ( : v.1985); 7ex (23 28.v.1991). : 1ex ( : 30.v.1985). : 1ex ( : 30.v.1970); 1ex (26.v.1985); 15ex (9 14.vi.1991). : 422, 4 8.(i- 2, iv- 4, v- 156, vi- 241, vii- 16, viii- 3ex). 101. Sericania latisulcata Murayama, 1941: 36 ( 9-87, 16-87, 22-87) Sericania latisulcata Murayama, Annot. Zool. Japan, 20(1): 36, 1941: Murayama, 1954: 69; Cho, 1969: 662 ( ); Kim, 1978: 383; Stebnicka, 1980: 206; Kim and Nam, 1982a: 155; Kim and Lee, 1991f: 49; Kim, 2001b: 59; Loebl and Smetana, 2006: 246. Serica latisulcata: ESK and KSAE, 1994: 151. : (Kim, 1978). : ( )... 4,. 9 4.5. 1.6, 5 1 4 3/5..,

: : 203, 1... 2. 1 ( ). Ad:,.,,.,. BL: 9 12 mm, BW: 5 6 mm. : Kim and Kim (2003c). :. :,,. : SK-GB, GS ( ),,, CB, JN,,. (8ex): [SWU] : 1 ( : 25.v.1995). [SNU] : 1ex ( : 1.v.1992). : 2ex ( : 23.vi.1993); 1ex ( : 23.vi.1995); 1ex ( : 23.vi.1995). [EWU] : 1ex ( : 23.v.1991); 1ex ( : 7.vi.1981). 102. Sericania koryoensis Murayama, 1935: 8 ( 9-88, 16-88, 22-88) Sericania koryoensis Murayama, J. Chosen Nat. Hist. Soc., 20: 8, 1935: Murayama, 1938: 15; 1954: 71; Miwa and Chûjô, 1939: 59; Cho, 1957: 297; 1969: 662; Kim, 1981: 344; 2001: 56; Stebnicka, 1980: 206; Kim and Lee, 1991b: 67; 1991f: 49; Kim et al., 2004: 117; Loebl and Smetana, 2006: 246. Serica koryoensis: ESK and KSAE, 1994: 151. : (Cho, 1969). : (Koryo= 1 ).,.,. 10 4, 1.6, 3..,,, 1., 1. 2. 1. Ad:, 0.8 mm.,.. BL: 8 10 mm, BW: 4.5 5.5 mm. : Kim and Kim (2003c).

204 :. :,. : SK-GW,,,,,, GG,,,,,,,, CB,, GBN,,,,,,, JBN,,,. (105ex): [SWU] : 17ex (,,,,, : v vi.1974 1999). : 8ex ( : iv vi.1991 2000); 4ex ( : v.1962 1982). : 1ex ( : 17.vi.2003); 2ex (, : v.1984, 1987). : 26ex (,,. : v vi.1978 1996); 1ex ( : 1.v.2006). : 2ex (, : iv, v.1966, 1968). : 13ex ( : v.1983, 1993). : 1ex ( : iv.1990). [NAAS] : 1ex ( : 19.iv.1983). [SNU] : 2ex ( : 12.v.1992); 2ex ( : v.1982, 1983). : 5ex ( : v.1991 1996); 1ex ( : 19.vi.1992); 1ex ( : v.1991). : 1ex ( : 12.v.1992). : 1ex ( : 10.v.1992). [KU] : 2ex ( : v, vi.1997, 1999); 2ex ( : vi.1992). : 1ex ( : 10.v.1997); 1ex ( : 19.v.1973). : 2ex ( : vi.1989). : 2ex ( : v.1983). : 2ex ( : v.1988). [EWU] : 2ex ( : 10.v.1990). : 1ex ( : 24.v.1987). : 1ex ( : 6.v.1977). 103. Sericania yamauchii Sawada, 1938: 8 ( 9-89, 16-89, 22-89) Sericania yamauchii Sawada, Nippon no Kŭchŭ, II(1): 8, 14, 1938: Kim and Lee, 1991f: 51; Kim, 1992: 105; 1993: 65; 1995a: 164; 2001b: 57; Loebl and Smetana, 2006: 246. Serica yamauchii: ESK and KSAE, 1994: 151. Sericania hasegawai Murayama, Trans. Biol. Soc. Machoukuo. 4 (1): 21, 1941 (, 1 ; Yablonia, 1 ): 1954: 72; Cho, 1969: 662 ( ); Kim, 1978: 383; Stebnicka, 1980: 206. : (Hiko, Kiushiu). (S. koryoensis). 1.,. 9 4 1.6, 3..,,, 1.. 2. 1. Ad: 1.2 mm,,.,,

: : 205 85.,.. BL: 9.5 12 mm, BW: 5 6 mm. : Kim and Kim (2003c). :,,. :,,. : SK-GW,,,,, GG,,,,,,, CBN,, GB, JBN,,,,, JJ. (35ex): [SWU] : 1ex ( : 4.vi.1999); 2ex (, : v, vii.1983, 1995); 1ex ( : v.1970). : 4ex (,, : iv vi.1961 1999). : 1ex ( : 4.vi.2000); 3ex ( : iv.1983). : 8ex ( : vi.1981); 1ex ( : 4.v.1991). : 1ex ( : v.1999). [SNU] : 1ex ( : 29.v.1997); 1ex ( : 23.vi.1995); 1ex ( : 24.vi.1995). [KU] : 1ex ( : 8.vi.1995). : 1ex ( : 4.vi.2000). : 1ex ( : 6.vi.1981). [EWU] : 2ex ( : 27.v.1973, 1.vi. 1991). [HMNH-NK] : 1ex ( : 7.vi.1991). : 1ex ( : 22.v.1985); 2ex (25.v.1991). : 1ex ( : 12.vi.1991). Sericania (= ) 1. Sericania angulata Lewis, 1985 Sericania angulata Lewis, Ann. Mag. Nat. Hist., 6(XVI): 392, 1895: Lee et al., 1985: 125; Kim and Lee, 1991f: 51; ESK and KSAE, 1994: 151 (Serica, ),. 2. Sericania imadatei Sawada, 1955 Sericania imadatei Sawada, J. Agr. Sci. Tokyo Nogyo Daigaku, 2: 569, 1955: Kim et al., 1974a: 229 ( ); Lee et al., 1981: 82 ( ); Kim and Lee, 1991f: 50; ESK and KSAE, 1994: 151 (Serica, ),.

206 3. Sericania kamiyai Sawada, 1938 Sericania kamiyai Sawada, Nippon no Kochu, 2(1): 12,18, 1938: Kim and Kim, 1972a ( ): 83; Kim and Lee, 1991f: 50; Serica: ESK and KSAE, 1994: 151 ( ), Nipponoserica opacicarina. Genus Nipponoserica Nomura, 1973: 120. Pseudomaladera Nikolayev, 1980: 40 ( : Serica koltzei Reitter, 1897 )., 9 3...,, ( ).. : Serica similis Lewis, 1895: 391. : 19 (Loebl and Smetana, 2006) ( 2 ). :. 1. 8 mm.. elliptica 8.5 mm.. opacicarina 104. Nipponoserica elliptica (Murayama), 1938: 17 ( 9-90, 16-90, 22-90) Serica elliptica Murayama, Annot. Zool. Japan, 17(1): 17, 1938: Murayama, 1954: 32; Cho, 1963: 217; 1969: 650 ( ); Cho et al., 1968: 264; Kim and Nam, 1982a: 154; Kim and Lee, 1991b: 67; 1991f: 55. Nipponoserica elliptica: Kim, 2001b: 61; Kim and Kim, 2003: 76; Loebl and Smetana, 2006: 240.

: : 207 : (Cho, 1963; Cho et al., 1968). : (, Koryo= ).,.,,. 9 3 1.2 1.4,..,.,, 1.., 2/3.. Ad:.,.. BL: 8 11 mm, BW: 5 6.5 mm. : Kim and Kim (2003c). :. :,,. : SK-GW, GS-,,,,,, CB, GBN,,,,,, JBN,,,,, JJ. (40ex): [SWU] : 1ex ( : 27.ix.2000); 16ex (, : 24.viii); 1ex (v.1987); 1 ( : 7.vii.1998); 1 ( : 2.vi.2001); 1ex ( : 18.vii.2002). : 5ex (,,,, : v viii.1974 1992); 1ex ( : 18.vi.1998); 1 ( : 11 17.v.1991); 1ex ( : 19.v.2003). : 3ex (,, : v.1992, 1994); 1 ( : 25 26.vii.1990). [SNU] : 1ex ( : 24.vii.1974). : 1ex ( : 15.vii.1976); 1ex ( : 9.vii.1969). [EWU] : 2ex (, : 5.vi.1971); 2ex ( : vi.1976, 1990). 105. Nipponoserica opacicarina Kim et Kim, 2003: 76 ( 9-91, 16-91, 22-91) Nipponoserica opacicarina Kim et Kim, Ins. Koreana, 20(1): 76, 2003: Loebl and Smetana, 2006: 240. : Serica brunnea (Linné), Faun. Suec.: 138, 1758 (Scarabaeus brunneus): Murayama, 1938: 9; 1954: 30; Cho, 1969: 650 ( ); Stebnicka, 1980: 206; Kim and Nam, 1982a: 154; Kim and Lee, 1991b: 67; 1991f: 54; ESK and KSAE, 1994: 151; Kim, 1995a: 164; 2001: 62. Sericania kamiyai (Sawada), Nippon-no-Kochu, 1 (1): 27, 1937 (Serica): Kim and Kim, 1972a: 83. Maladera kamiyai: Kim and Chang, 1987a: 104.

208 :.,.., 9 2,..,.,, 1..,.,, 3/4. 1 ( ). Ad: (N. elliptica)... BL: 6.5 8.5 mm, BW: 4 5 mm. : Kim and Kim (2003c). :. :,. :. (22ex): [SWU] : 1 [ : 22.v.1992 ( )]. : 1 ( : 26.vi.1984). : 1 ( : 23.v.1984); 1 ( : 23.v.1988). : 1 ( : 10.viii.1987); 1 ( : 24.vii.1986). : 1 ( : 24.v.1994)- ; : 1ex ( : 19.v.1935). : 1 ( : 7.vi.1974). : 1ex ( : 1.vi.1984); 1 ( : 7.v.1984); 1 1 (1 22.vi.1984); 1 ( : 3.vi.1992). : 1 ( : 11.v.1992); 1 ( : 21.v.1992). [NAAS] : 2ex ( : v, vi.1925, 1926); 1ex ( : 6.i.1922). [SNU] : 1ex ( : 28.v.1990); 1ex ( : 24.v.1990). [EWU] : 1ex ( : 13.vi.1984). : Murayama (1954) Serica brunnea Nomura (1973). (Kim and Kim, 2003) London. Nipponoserica (= ) 1. Nipponoserica similis (Lewis), 1895 Serica similis Lewis, Ann. Mag. Nat. Hist., 6(XVI): 391, 1895 (Japan): Okamoto, 1924: 172; KZS, 1968: 136 ( ) 17; Nipponoserica-Stebnicka, 1980: 250; Kim and Lee, 1991f: 51 Murayama

: : 209 (1954) Okamoto. Genus Serica MacLeay, 1819: 146. Trichoserica Reitter, 1896: 181 ( : Trichoserica fulvopubens Reitter, 1896, ). Ophthalmoserica Brenske, 1897: 356 ( : Serica thibetana Brenske, 1897; Ahrens 1999c ). Podoserica Breit, 1912: 202 ( : Podoserica reitteri Breit, 1912, monotypy).,. 9 3.. 2, 2..,.,.. Serica, Trichoserica, Paraserica, Omaloplia, Ophthalmoserica, Maladera 11 4. : Scarabaeus brunneus (Linné) 1758. : 300 (?) ( )( 170 )( 4 ). :. : Brenske (1897), Reitter (1902), Murayama (1954), Kim et al. (1991f, 1997b, 2001b, 2003). 1. septentrionalis 2 2.. 2 hirsuta. 2 3 3.,. Ad: fulvopubens. Ad: lutea

210 106. ( ) Serica fulvopubens (Reitter), 1986: 184 ( 9-92, 16-92, 22-92) Trichoserica fulvopubens (Reitter), Entom. Zeit., 15: 184, 1986. Omalopia polita Gebler, Nouv. Mem. Soc. Nat. Moscou, II: 53, 1832. Serica polita: Murayama, 1935: 2; 1937: 32; 1938: 7; 1937b: 15; 1941: 17; 1954: 23; Sawada, 1937: 9; Miwa and Chûjô, 1939: 53; Cho, 1957: 295; 1969: 649 ( ); Kim and Kim, 1972a: 83; Nomura, 1972: 111; Kim et al., 2002: 120; Loebl and Smetana, 206: 244. Trichoserica polita: Stebnicka, 1980: 250 (=S. hirsuta?); Kim and Lee, 1991b: 67; 1991f: 52; Kim et al., 1998: 170; 1999: 129; Park et al., 1993: 178; ESK and KSAE, 1994: 151; Kim, 1992: 105; 1993: 65; 1995a: 164; 2001b: 62. Serica fulvopubens: Nikolayev, 2002: 120.,..,,., 9 2,...,,.., 2/3. 1 ( ) 2. Ad: 2.,. BL: 8.5 10 mm, BW: 4.5 5.5 mm. : Kim and Kim (2003c). :,,,. :,,,. : NK-HB,,, YG, HN,,, PB, WB, GG, SK-GW,,,,,,,,,, GS,,,,,,,,,,,,,,, GBN,,,,,,,,, JBN,,,,,,,, JJ-. (172ex): [SWU] : 21ex (,,,,, : iv ix.1958 2000); 1ex ( : 19.viii.2001). : 20ex (,,,,, : v vii.1958 1994). : 1ex ( : vii.1987). : 11ex ( : 29.vi.1998); 18ex (,,,,,, : v ix.1974 1999). : 21ex (,,,,, : v viii.1958 1986). : 1ex (vii.1974). [NAAS] : 1ex [Gesseji ( ): 8.vii.1923]. [SNU] : 3ex ( : vi.1988); 1ex ( : 1997); 1ex ( : vi.1988). : 1ex ( : vi.1984). : 17ex ( : vii.1976); 1ex

: : 211 ( : vi.1997); 2ex ( : vi, vii.1988, 1991). : 1ex (vii.1971). [KU] : 2ex (, : vi.1992, 1999). : 9ex (,,, : vi.1977 1989). : 8ex ( : vi.2000). : 1ex ( : vi.1997). : 4ex (,, : iv viii.1989 1996). : 14ex ( : vi.1977 1998). [EWU] : 1ex ( : 31.v.1978); 1ex ( : 7.vi.1975); 1ex ( : 9.vi.1974); 1ex ( : 9.vi.1984); 1ex ( : 28.v.1988). : 1ex ( : 29.vii.1983). [HMNH-NK] : 3ex ( : 18 19.vii.1977); 2ex ( : 27.vii.1975); 2ex ( : 22.vii.1975). : 134, 4 9. (iv- 2, v- 10, vi- 87, vii- 21, viii- 7, ix- 8ex). :. 107. Serica lutea Kim and Kim, 2003: 75 Serica lutea Kim and Kim, Ins. Koreana, 20(1): 75, 2003. Trichoserica boops Waterhouse, Trans. Ent. Soc. Lond.: 101, 1875 ( )( ): Kim and Lee, 1991f: 53. :.,.,. 2. 9, 2,.,.,.,.,.,, (, spurs). Ad:,..,. BL: 6 9.5 mm, BW: 3 5 mm. :. :. (2 types): [SWU] : 1 [ : 29.v.1975 ( )]; 1 [ : 18.vi.1978 ( )]. : (Kim and Lee, 1991f) Trichoserica boops.

212 108. Serica septentrionalis Murayama, 1935: 7 ( 9-93, 16-93, 22-93) Serica septentrionalis Murayama, Journ. Chosen Nat. Hist. Soc., 20: 7, 1935: Murayama, 1937a: 33; 1938a: 9; 1954: 29; Sawada, 1937a: 9; Miwa and Chûjô, 1939: 45; Cho, 1957: 295; 1969: 650 ( ); Stebnicka, 1980: 206; Kim, 2001b: 64; Loebl and Smetana, 206: 245. Trichoserica septentrionalis: Kim and Lee, 1991f: 53; ESK and KSAE, 1994: 152. : ( HB).,.,.. 9, 2..,.,,..,,. Ad:,. BL: 7 8 mm, BW: 4 4.5 mm. :. :. : NK-HB: 1 1 ( : 20.vii.1934);?: 1 [ : 16.vii.1934 (Murayama, 1954)]. :. 109. Serica hirsuta Kim and Kim, 2003: 74 ( 9-94, 16-94, 22-94) Serica hirsuta Kim and Kim, Ins. Koreana, 20(1): 74, 2003: Loebl and Smetana, 206: 244. :.,.. 2,. 1/2.,.,,.,.

: : 213,, (, spurs). Ad:,,. BL: 7.3 9.8 mm, BW: 3.8 5 mm. : Kim and Kim (2003c). :. :,. (53ex): [SWU] : 1 1 [ : 26.vi.1996 ( )]; 4 (24 25.vi.1996); 1 ( : 22.iv.1984); 1 ( : 21.v.1998); 1 ( : 19.vi.1974). : 4 ( : 5.vi.1977); 1 (29.v.1983); 1 (30.vi.1984). : 1 ( : 23.vii.1974); 2 (5 6. vi.1981); 1 ( : 24.v.1997). : 1 ( : 4.vi.1986). : 2 ( : 19.vi. 1982); 1 3 (23.vi.1987); 1 (21.v.1998)- ; : 1 ( : 6.vi.1974). : 1ex ( : 24.vi.1991); 1 1 ( : 28.v.1982); 1 (31.v.1986); 1 ( : 8.vi.1968); 1 (2.vi.1984); 2 ( : 2.vi.1985); 1 ( : 23.vi.1974); 1 ( : 12.vi.1978); 1 ( : 5.v.1994); 1 ( : 19.vi.1982); 1 ( : 15.v.1994). : 1 ( : 10.vii.1977); 1 (26.v.1996); 1 ( : 4.vi.1978); 1 ( : 5.iv.1989); 1 ( : 23.v.1989); 1 ( : 29.vi.1998). : 1 ( : 9.vi.1972). : 1 ( : 24.vi.1986); 1 (16.vii.1984); 2ex ( : 13.vi.1997, 16.vii. 1976); 1 ( : 22.vi.1987). Serica (= ) 1. Serica boops Waterhouse, 1875 Serica boops Waterhouse, Trans. Ent. Soc., Lond.: 101, 1875 ( ): Okamoto, 1924: 172 (Saishuto=, 1 ); Cho, 1963: 217 ( ); 1969: 649 ( ), 8; Ophthalmoserica: Hoikusha, 1956: 99; Trichoserica: Kim and Lee, 1991f: 53 Murayama (1937a) Okamoto,. 2. Serica brunnea (Linné), 1758 Scarabaeus brunnea Linné, Syst. Nat., ed. 10: 352, 1758: Serica: Murayama, 1938: 9; 1941: 18; 1954: 30; Kim and Nam, 1982a: 154 ( ) 4 Nipponoserica opacicarina.

214 3. Serica formosana Moser, 1915 Serica formosana Moser, 1915; Deut. ent. Zeit.: 338, 1915 (Taiwan); Kinoshita and Niijima, 1923: 238 ( ); ESK and KSAE ( ) Moser. 4. Serica sawadai Nomura, 1959 Ophthalmoserica sawadai Nomura, Toho-Gakuho, Honshu Japan, (9): 46, 1959 (Taiwan): Nam and Kim, 1982: 129 (Mt. Jirisan).

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227 1 9 : (, ) 10 16 : 17 22 : (,, ) 1 1. Valgus koreanus Sawada (, ) 2. Nipponovalgus angusticollis (Waterhouse) ( ) 3. Osmoderma opicum Lewis ( ) 4. Gnorimus subopacus Motschulsky 5. Trichius succinctus (Pallas) 6. Dicranocephalus adamsi Pascoe (, ) 7. Rhomborrhina polita Waterhouse 8. Pseudotorynorrhina japonica (Hope) 9. Cetonia pilifera pilifera (Motschulsky) 9a. Cetonia pilifera magnifica Ballion 10. Protaetia lugubris (Herbst) 11. Protaetia brevitarsis seulensis (Kolbe) 2 11. Protaetia brevitarsis seulensis (Kolbe) 12. Protaetia famelica scheini (Mikšić ) 13. Protaetia mandschuriensis (Schürhoff) 14. Protaetia orientalis submarmorea (Burmeister) 15. Anthracophora rusticola Burmister 16. Glycyphana fulvistemma Motschulsky 17. Gametis jucunda (Faldermann) 18. Clinterocera obsoleta (Fairmaire) 19. Adoretus tenuimaculatus Waterhouse 20. Adoretus hirsutus Ohaus 3 21. Parastasia ferrieri Nonfried 22. Popillia flavosellata Fairmaire 23. Popillia mutans Newman 24. Popillia quadriguttata (Fabricius)

228 25. Popillia formosana Arrow 26. Proagopertha lucidula (Faldermann) 27. Phyllopertha diversa Waterhouse (,, ) 28. Mimela fusania Bates 29. Mimela splendens (Gyllenhal) 30. Mimela holosericea (Fabricius) 4 31. Mimela testaceipes (Motschulsky) 32. Spilota plagiicollis (Fairmaire) 33. Blitopertha conspurcata (Harold) (, ) 34. Blitopertha pallidipennis Reitter (,,, ) 35. Blitopertha orientalis (Waterhouse) (, ) 36. Anomala sieversi Heyden 37. Anomala albopilosa Hope 38. Anomala japonica Arrow 5 39. Anomala mongolica (Faldermann) 40. Anomala chamaeleon Fairmaire 41. Anomala rufocuprea Motschulsky 42. Anomala koreana Kim 43. Anomala luculenta Erichson 44. Anomala corpulenta Motschulsky 45. Bifurcanomala aulax (Wiedemann) 46. Chejuanomala quelparta (Okamoto) 47. Allomyrina dichotoma (Linné) (, ) 48. Eophileurus chinensis (Faldermann) ( ) 6 48. Eophileurus chinensis (Faldermann) ( ) 49. Pentodon quadridens (Gebler) 50. Ectinohoplia rufipes (Motschulsky) 51. Hoplia (Euchromoplia) aureola (Pallas) 52. Apogonia cribricollis Burmeister 53. Apogonia cupreoviridis Kolbe 54. Brahmina rubetra faldermanni (Murayama) 55. Lasiopsis sahlbergi (Mannerheim)

229 56. Pseudosymmchia impressifrons (Fairmaire) 57. Sophrops heydeni (Brenske) 58. Sophrops striata (Brenske) 59. Holotrichia niponensis (Lewis) 60. Holotrichia parallela (Motschulsky) 7 60. Holotrichia parallela (Motschulsky) 61. Holotrichia koraiensis Murayama 62. Holotrichia picea (Waterhouse) 63. Holotrichia ernesti Reitter 64. Holotrichia diomphalia (Bates) 65. Holotrichia oblita (Faldermann) 66. Holotrichia reticulata Murayama 67. Heptophylla picea Motschulsky 68. Hilyotrogus bicoloreus (Heyden) 69. Polyphylla laticollis manchurica Semenov (, ) 70. Melolontha incana (Motschulsky) (, ) 71. Melolontha insulana (Moser) (, ) 8 72. Gastroserica herzi (Heyden) 73. Maladera holosericea (Scopoli) 74. Maladera verticalis (Fairmaire) 75. Maladera castanea koreana Kim and Kim 76. Maladera ovatula (Fairmaire) 77. Maladera aureola (Murayama) 78. Maladera renardi (Ballion) 79. Maladera schoenfeldti (Murayama) 80. Maladera gibbiventris (Brenske) 81. Maladera infuscata (Moser) 82. Maladera fusania (Murayama) 83. Maladera opaciventris (Moser) 84. Maladera orientalis (Motschulsky) 9 85. Maladera cariniceps (Moser) 86. Sericania fuscolineata Motschulsky (, )

230 87. Sericania latisulcata Murayama 88. Sericania koryoensis Murayama 89. Sericania yamauchii Sawada 90. Nipponoserica elliptica (Murayama) 91. Nipponoserica koltzei (Reitter) 92. Serica fulvopubens (Reitter) (, ) 93. Serica septentrionalis Murayama 94. Serica hirsuta Kim and Kim 10 1. Valgus koreanus Sawada (, ) 2. Nipponovalgus angusticollis (Waterhouse) ( ) 3. Osmoderma opicum Lewis ( ) 4. Gnorimus subopacus Motschulsky 5. Trichius succinctus (Pallas) 6. Dicranocephalus adamsi Pascoe (, ) 7. Rhomborrhina polita Waterhouse 8. Pseudotorynorrhina japonica (Hope) 9. Cetonia pilifera pilifera (Motschulsky) 9a. Cetonia pilifera magnifica Ballion 10. Protaetia lugubris (Herbst) 11. Protaetia brevitarsis seulensis (Kolbe) 12. Protaetia famelica scheini (Mikšić ) 13. Protaetia mandschuriensis (Schürhoff) 11 14. Protaetia orientalis submarmorea (Burmeister) 15. Anthracophora rusticola Burmister 16. Glycyphana fulvistemma Motschulsky 17. Gametis jucunda (Faldermann) 18. Clinterocera obsoleta (Fairmaire) 19. Adoretus tenuimaculatus Waterhouse 20. Adoretus hirsutus Ohaus 21. Parastasia ferrieri Nonfried 22. Popillia flavosellata Fairmaire 23. Popillia mutans Newman 24. Popillia quadriguttata (Fabricius) 25. Popillia formosana Arrow

231 26. Proagopertha lucidula (Faldermann) 27. Phyllopertha diversa Waterhouse (, ) 12 27. Phyllopertha diversa Waterhouse ( ) 28. Mimela fusania Bates 29. Mimela splendens (Gyllenhal) 30. Mimela holosericea (Fabricius) 31. Mimela testaceipes (Motschulsky) 32. Spilota plagiicollis (Fairmaire) 33. Blitopertha conspurcata (Harold) (, ) 34. Blitopertha pallidipennis Reitter (,,, ) 35. Blitopertha orientalis (Waterhouse) (, ) 36. Anomala sieversi Heyden 37. Anomala albopilosa Hope 13 38. Anomala japonica Arrow 39. Anomala mongolica (Faldermann) 40. Anomala chamaeleon Fairmaire 41. Anomala rufocuprea Motschulsky 42. Anomala koreana Kim 43. Anomala luculenta Erichson 44. Anomala corpulenta Motschulsky 45. Bifurcanomala aulax (Wiedemann) 46. Chejuanomala quelparta (Okamoto) 47. Allomyrina dichotoma (Linné) (, ) 48. Eophileurus chinensis (Faldermann) (, ) 49. Pentodon quadridens (Gebler) 50. Ectinohoplia rufipes (Motschulsky) 14 51. Hoplia (Euchromoplia) aureola (Pallas) 52. Apogonia cribricollis Burmeister 53. Apogonia cupreoviridis Kolbe 54. Brahmina rubetra faldermanni (Murayama) 55. Lasiopsis sahlbergi (Mannerheim) 56. Pseudosymmchia impressifrons (Fairmaire)

232 57. Sophrops heydeni (Brenske) 58. Sophrops striata (Brenske) 59. Holotrichia niponensis (Lewis) 60. Holotrichia parallela (Motschulsky) 61. Holotrichia koraiensis Murayama 62. Holotrichia picea (Waterhouse) 63. Holotrichia ernesti Reitter 64. Holotrichia diomphalia (Bates) 65. Holotrichia oblita (Faldermann) 66. Holotrichia reticulata Murayama 67. Heptophylla picea Motschulsky 68. Hilyotrogus bicoloreus (Heyden) 15 69. Polyphylla laticollis manchurica Semenov (, ) 70. Melolontha incana (Motschulsky) (, ) 71. Melolontha insulana (Moser) (, ) 72. Gastroserica herzi (Heyden) 73. Maladera holosericea (Scopoli) 74. Maladera verticalis (Fairmaire) 75. Maladera castanea koreana Kim and Kim 76. Maladera ovatula (Fairmaire) 77. Maladera aureola (Murayama) 78. Maladera renardi (Ballion) 79. Maladera schoenfeldti (Murayama) 80. Maladera gibbiventris (Brenske) 81. Maladera infuscata (Moser) 82. Maladera fusania (Murayama) 83. Maladera opaciventris (Moser) 16 84. Maladera orientalis (Motschulsky) 85. Maladera cariniceps (Moser) 86. Sericania fuscolineata Motschulsky (, ) 87. Sericania latisulcata Murayama 88. Sericania koryoensis Murayama 89. Sericania yamauchii Sawada 90. Nipponoserica elliptica (Murayama)

233 91. Nipponoserica koltzei (Reitter) 92. Serica fulvopubens (Reitter) (, ) 93. Serica septentrionalis Murayama 94. Serica hirsuta Kim and Kim 17 1. Valgus koreanus Sawada 2. Nipponovalgus angusticollis (Waterhouse) 4. Gnorimus subopacus Motschulsky 5. Trichius succinctus (Pallas) 6. Dicranocephalus adamsi Pascoe 7. Rhomborrhina polita Waterhouse 8. Pseudotorynorrhina japonica (Hope) 9. Cetonia pilifera pilifera (Motschulsky) 9a. Cetonia pilifera magnifica Ballion 10. Protaetia lugubris (Herbst) 11. Protaetia brevitarsis seulensis (Kolbe) 12. Protaetia famelica scheini (Mikšić ) 13. Protaetia mandschuriensis (Schürhoff) 14. Protaetia orientalis submarmorea (Burmeister) 15. Anthracophora rusticola Burmister 16. Glycyphana fulvistemma Motschulsky 17. Gametis jucunda (Faldermann) 18. Clinterocera obsoleta (Fairmaire) 19. Adoretus tenuimaculatus Waterhouse 20. Adoretus hirsutus Ohaus 18 21. Parastasia ferrieri Nonfried 22. Popillia flavosellata Fairmaire 23. Popillia mutans Newman 24. Popillia quadriguttata (Fabricius) 25. Popillia formosana Arrow 26. Proagopertha lucidula (Faldermann) 27. Phyllopertha diversa Waterhouse 28. Mimela fusania Bates 29. Mimela splendens (Gyllenhal) 30. Mimela holosericea (Fabricius)

234 31. Mimela testaceipes (Motschulsky) 32. Spilota plagiicollis (Fairmaire) 33. Blitopertha conspurcata (Harold) 34. Blitopertha pallidipennis Reitter 35. Blitopertha orientalis (Waterhouse) 36. Anomala sieversi Heyden 37. Anomala albopilosa Hope 38. Anomala japonica Arrow 19 39. Anomala mongolica (Faldermann) 40. Anomala chamaeleon Fairmaire 41. Anomala rufocuprea Motschulsky 42. Anomala koreana Kim 43. Anomala luculenta Erichson 44. Anomala corpulenta Motschulsky 45. Bifurcanomala aulax (Wiedemann) 46. Chejuanomala quelparta (Okamoto) 47. Allomyrina dichotoma (Linné) 48. Eophileurus chinensis (Faldermann) 49. Pentodon quadridens (Gebler) 50. Ectinohoplia rufipes (Motschulsky) 51. Hoplia (Euchromoplia) aureola (Pallas) 52. Apogonia cribricollis Burmeister 53. Apogonia cupreoviridis Kolbe 20 54. Brahmina rubetra faldermanni (Murayama) 55. Lasiopsis sahlbergi (Mannerheim) 56. Pseudosymmchia impressifrons (Fairmaire) 57. Sophrops heydeni (Brenske) 58. Sophrops striata (Brenske) 59. Holotrichia niponensis (Lewis) 60. Holotrichia parallela (Motschulsky) 60. Holotrichia parallela (Motschulsky) 61. Holotrichia koraiensis Murayama 62. Holotrichia picea (Waterhouse) 63. Holotrichia ernesti Reitter

235 64. Holotrichia diomphalia (Bates) 66. Holotrichia reticulata Murayama 67. Heptophylla picea Motschulsky 68. Hilyotrogus bicoloreus (Heyden) 69. Polyphylla laticollis manchurica Semenov 21 70. Melolontha incana (Motschulsky) 71. Melolontha insulana (Moser) 72. Gastroserica herzi (Heyden) 73. Maladera holosericea (Scopoli) 74. Maladera verticalis (Fairmaire) 75. Maladera castanea koreana Kim and Kim 76. Maladera ovatula (Fairmaire) 78. Maladera renardi (Ballion) 79. Maladera schoenfeldti (Murayama) 80. Maladera gibbiventris (Brenske) 81. Maladera infuscata (Moser) 82. Maladera fusania (Murayama) 83. Maladera opaciventris (Moser) 84. Maladera orientalis (Motschulsky) 85. Maladera cariniceps (Moser) 22 86. Sericania fuscolineata Motschulsky 87. Sericania latisulcata Murayama 88. Sericania koryoensis Murayama 89. Sericania yamauchii Sawada 90. Nipponoserica elliptica (Murayama) 91. Nipponoserica koltzei (Reitter) 92. Serica fulvopubens (Reitter) 93. Serica septentrionalis Murayama 94. Serica hirsuta Kim and Kim

236 1 1 1 2 5 3 4 6 6 7 8 8 9 9a 9a 10 11

237 2 11 12 12 13 14 14 15 16 17 17 17 17 18 19 20

238 3 21 22 22 23 24 25 26 27 27 27 28 28 29 30 30

239 4 31 31 32 33 33 34 34 34 34 35 35 36 36 37 38

240 5 39 40 40 41 42 42 43 43 44 45 45 46 47 47 48

241 6 48 50 50 50 49 51 51 52 53 54 55 56 57 58 59 60

242 7 60 61 62 63 64 65 66 67 68 69 69 70 70 71 71

243 8 72 72 72 73 74 75 76 77 78 79 80 81 82 83 84

244 9 85 86 86 87 88 88 89 90 91 92 92 93 94

245 10 1 1 2 3 4 5 6 6 7 8 8 9 9a 9a 10 11 11 12 12 13

246 11 14 14 15 16 17 17 17 17 18 19 20 21 22 22 23 24 25 26 27 27

247 12 27 28 28 29 30 30 31 31 32 33 33 34 34 34 34 35 35 36 36 37

248 13 38 39 40 40 41 42 43 43 44 45 45 46 47 47 48 48 49 50 50 50

249 14 51 51 52 53 54 55 56 57 58 59 60 60 61 62 63 64 65 66 67 68

250 15 69 69 70 70 71 71 72 72 72 73 74 75 76 77 78 79 80 81 82 83

251 16 84 85 86 86 87 88 88 89 90 91 92 92 93 94

252 17 1 2 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20

253 18 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38

254 19 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53

255 20 54 55 56 57 58 59 60 61 62 63 64 66 67 68 69

256 21 70 71 72 73 74 75 76 78 79 80 81 82 83 84 85

257 22 86 87 88 89 90 91 92 93 94